Heal for 40 every time you Heal an ally with Moonlight.
Level 4
Celestial Body
Armor
Increase your maximum Health by 250.
Level 5
Spirit Arrows
Moonlight
Reduce the Cooldown of Luna's redeploy by 0.2s after healing with Moonlight.
Level 2
Sky-Walker
Armor
Lower your gravity and increase your air control by 10% while firing your Light Bow.
Level 1
Crescent
Lunar Leap
Reset the Cooldown of Lunar Leap after dropping to or below 45% Health.
Level 3
Sacrifice
Guardian Spirit
[Guardian Spirit] If Io falls below 200 Health while Luna is active, she trades places with Luna, Heals for 50% of her Health, and Luna dies in her stead.