Hull Piercer

Harpoon

Reduce the Cooldown of Harpoon by 6s.

Level 5

Expansive Vault

Shortcut

Jump height is increased for your first jump within 1s after exiting a Shortcut.

Level 1

Dark Bargain

Shortcut

Heal for 200 every 1s while standing near a Shortcut.

Level 4

Heave Away

Broadside

Reduce the Cooldown of Harpoon by 0.8s for each enemy hit by Broadside.

Level 4

Blow the Man Down

Harpoon

Enemies hit by Harpoon’s Slow have their Movement Speed reduced by an additional 6%.

Level 1

Abyss Spike

Harpoon

Harpoon lodges into terrain and explodes when an enemy comes within 20 units, dealing 1000 damage in a 35-unit-radius area.

Level 1

Level 3

Level 5

Level 2

Level 2

Level 3

Level 1

Level 3

Level 4

Level 2

Level 4

Level 2

Level 1

Hull Piercer

Harpoon

Reduce the Cooldown of Harpoon by 6s.

Level 5

Expansive Vault

Shortcut

Jump height is increased for your first jump within 1s after exiting a Shortcut.

Level 1

Dark Bargain

Shortcut

Heal for 200 every 1s while standing near a Shortcut.

Level 4

Heave Away

Broadside

Reduce the Cooldown of Harpoon by 0.8s for each enemy hit by Broadside.

Level 4

Blow the Man Down

Harpoon

Enemies hit by Harpoon’s Slow have their Movement Speed reduced by an additional 6%.

Level 1

Abyss Spike

Harpoon

Harpoon lodges into terrain and explodes when an enemy comes within 20 units, dealing 1000 damage in a 35-unit-radius area.

Level 1

Level 3

Level 4

Level 2

Level 4

Level 2

Level 1

Level 3

Level 4

Level 2

Level 4

Level 2

Level 1

Broken Deity

Guardian Spirit

Increase Luna's deploy range by 20%.

Level 1

Sky-Walker

Armor

Lower your gravity and increase your air control by 30% while firing your Light Bow.

Level 3

Half Moon

Lunar Leap

Reduce the Cooldown of Lunar Leap by 4s.

Level 5

Full Moon

Lunar Leap

Gain a 300-Health Shield for 3s after activating Lunar Leap.

Level 3

Crescent

Lunar Leap

Reset the Cooldown of Lunar Leap after dropping to or below 45% Health.

Level 3

Sacrifice

Guardian Spirit

[Guardian Spirit] If Io falls below 200 Health while Luna is active, she trades places with Luna, Heals for 50% of her Health, and Luna dies in her stead.

Level 1

The Great Worm

Form Swap

Gain a 240-Health Shield for 2s after swapping into Travel Form.

Level 3

Violent Birth

Hardening

Reduce the Cooldown of Acceleration by 1.6s after activating Hardening.

Level 2

Triumphant Return

Form Swap

Reduce your acid consumption by 90% for 0.8s after swapping into Planted Form.

Level 2

Deadly Pursuit

Form Swap

Increase your Movement Speed by 20% for 3s after swapping into Travel Form.

Level 3

Towering Presence

Armor

Increase the Health of your Ultimate Health pool by 1000.

Level 5

Corrosive Acid

Caustic Spray

Augment your acid, increasing the damage done by Caustic Spray to 42, the damage of acid pools to 40, and increase the Movement Speed of allies within your acid pools by 20%.

Level 1

Broken Deity

Guardian Spirit

Increase Luna's deploy range by 20%.

Level 1

Sky-Walker

Armor

Lower your gravity and increase your air control by 30% while firing your Light Bow.

Level 3

Half Moon

Lunar Leap

Reduce the Cooldown of Lunar Leap by 4s.

Level 5

Full Moon

Lunar Leap

Gain a 300-Health Shield for 3s after activating Lunar Leap.

Level 3

Crescent

Lunar Leap

Reset the Cooldown of Lunar Leap after dropping to or below 45% Health.

Level 3

Sacrifice

Guardian Spirit

[Guardian Spirit] If Io falls below 200 Health while Luna is active, she trades places with Luna, Heals for 50% of her Health, and Luna dies in her stead.

Level 1

Follow the Scent

Fire Spit

Increase the size of your Fire Spit by 48%.

Level 2

Rain of Fire

Thrust

Consume 30% less Booster fuel for 5s after using Thrust.

Level 2

Condescension

Fire Spit

Reduce the Cooldown of Thrust by 2.4s for each enemy hit with Fire Spit.

Level 2

Masterful

Fire Spit

Reduce the Cooldown of Fire Spit by 2.5s for each enemy hit by it.

Level 5

Lung Capacity

Fire Spit

Heal for 200 for each enemy hit with Fire Spit.

Level 4

Combustible

Fire Spit

Fire Spit applies a Knockback and deals an additional 300 damage over 2s.

Level 1

Hull Piercer

Harpoon

Reduce the Cooldown of Harpoon by 6s.

Level 5

Expansive Vault

Shortcut

Jump height is increased for your first jump within 1s after exiting a Shortcut.

Level 1

Dark Bargain

Shortcut

Heal for 200 every 1s while standing near a Shortcut.

Level 4

Heave Away

Broadside

Reduce the Cooldown of Harpoon by 0.8s for each enemy hit by Broadside.

Level 4

Blow the Man Down

Harpoon

Enemies hit by Harpoon’s Slow have their Movement Speed reduced by an additional 6%.

Level 1

Abyss Spike

Harpoon

Harpoon lodges into terrain and explodes when an enemy comes within 20 units, dealing 1000 damage in a 35-unit-radius area.

Level 1

Hull Piercer

Harpoon

Reduce the Cooldown of Harpoon by 6s.

Level 5

Expansive Vault

Shortcut

Jump height is increased for your first jump within 1s after exiting a Shortcut.

Level 1

Dark Bargain

Shortcut

Heal for 200 every 1s while standing near a Shortcut.

Level 4

Heave Away

Broadside

Reduce the Cooldown of Harpoon by 0.8s for each enemy hit by Broadside.

Level 4

Blow the Man Down

Harpoon

Enemies hit by Harpoon’s Slow have their Movement Speed reduced by an additional 6%.

Level 1

Abyss Spike

Harpoon

Harpoon lodges into terrain and explodes when an enemy comes within 20 units, dealing 1000 damage in a 35-unit-radius area.

Level 1

Swift Strikes

Deadly Momentum

Increase the travel distance of Deadly Momentum by 20%.

Level 4

Rogues of a Feather

Love

Heal for 30 after hitting an enemy with Love.

Level 2

Both Sides Of The Coin

Weapon

Heal for 45 after hitting an enemy with War.

Level 3

Spin to Win

Deadly Momentum

Increase the size of the reactivation whirl of Deadly Momentum by 16%.

Level 2

Deadly Flourish

Deadly Momentum

Increase the travel speed of Deadly Momentum by 32%.

Level 4

Everywhere at Once

The Cooldown of Deadly Momentum is permanently set to 5s and cannot be modified, reduced, or increased through other means. Gain 1 stack of Piercing and Sharp Momentum for each enemy hit with Deadly Momentum.

Level 1