Regenerate 4% Moonlight every 1s while within 30 units of Luna and not using Moonlight.
Level 2
Sky-Walker
Armor
Lower your gravity and increase your air control by 10% while firing your Light Bow.
Level 1
Celestial Body
Armor
Increase your maximum Health by 250.
Level 5
Sacrifice
Guardian Spirit
[Guardian Spirit] If Io falls below 200 Health while Luna is active, she trades places with Luna, Heals for 50% of her Health, and Luna dies in her stead.