Extermination

Armor

Reduce the duration and effectiveness of Crowd Control by 15%.

Level 3

Deadly Predator

Weapon

Increase your maximum acid capacity by 8%.

Level 1

Raze and Burn

Piercing Quills

Heal for 50 after hitting an enemy with Piercing Quills.

Level 1

The Great Worm

Form Swap

Gain a 400-Health Shield for 2s after swapping into Travel Form.

Level 5

Towering Presence

Armor

Increase the Health of your Ultimate Health pool by 1000.

Level 5

Sight Begets Strength

Primal Vision

Allied players within range of your Primal Vision gain a stackable 90-Health Shield and 15% increased Movement Speed for 4s each time you pulse, but your Primal Vision has its Cooldown increased by 2s.

Level 1

Lorentz Configuration

Weapon

Increase your maximum Ammo by 30%.

Level 3

Retrograde

Astral Mark

Reduce the Cooldown of Astral Mark by 1.6s.

Level 4

Space Jam

Stellar Wind

Increase your jump height by 24% while using Stellar Wind.

Level 2

Solar Sails

Stellar Wind

Regenerate 4 Ammo every 1s while Stellar Wind is active.

Level 2

Penumbra

Weapon

Gain 20% Lifesteal.

Level 4

Binary Star

Weapon

Star Splitter now deals 360 damage every 0.4s, but reduces your maximum Ammo to 15.

Level 1

Octoppressor

Oppressor Mine

Increase the number of targets affected by Oppressor Mines by 4.

Level 4

Generator

Oppressor Mine

Reduce the Cooldown of Oppressor Mine by 2.4s.

Level 2

Prodigy

Sniper Mode

Increase your Movement Speed while in Sniper Mode by 25%.

Level 1

Quick Scope

Sniper Mode

Reduce scope-in and scope-out time by 60%.

Level 3

Amplitude

Oppressor Mine

Increase the range of Oppressor Mine beams by 100%.

Level 5

Oppression

Oppressor Mine

You can have two additional Oppressor Mines out at one time and you throw out two Mines per use of the ability instead of one, but your Mines have a lifetime of 6s and the Cooldown of Oppressor Mine is increased by 2s.

Level 1

Dark Sight

Rend Soul

Every Soul Charge you detonate generates 2 Ammo.

Level 2

Bane

Rend Soul

Every Soul Orb stack you detonate increases your Movement Speed by 3% for 4s, up to a maximum of 15%.

Level 3

Wickedness

Rend Soul

Reduce the Cooldown of Rend Soul by 1.0499999999999998s.

Level 3

Umbral Gait

Weapon

Your Movement Speed is increased by 24% while any enemy is marked with a Soul Orb stack.

Level 3

Sorceress

Weapon

Increase your Ultimate charge rate by 28%.

Level 4

Agony

Restore Soul

For each stack of Soul Orb detonated by Rend Soul, your next Restore Soul ignores 5% of Antihealing (excluding Healing disable effects) up to 25% (5 stacks) for up to 10s.

Level 1

The Great Worm

Form Swap

Gain a 240-Health Shield for 2s after swapping into Travel Form.

Level 3

Violent Birth

Hardening

Reduce the Cooldown of Acceleration by 1.6s after activating Hardening.

Level 2

Triumphant Return

Form Swap

Reduce your acid consumption by 90% for 0.8s after swapping into Planted Form.

Level 2

Deadly Pursuit

Form Swap

Increase your Movement Speed by 20% for 3s after swapping into Travel Form.

Level 3

Towering Presence

Armor

Increase the Health of your Ultimate Health pool by 1000.

Level 5

Corrosive Acid

Caustic Spray

Augment your acid, increasing the damage done by Caustic Spray to 42, the damage of acid pools to 40, and increase the Movement Speed of allies within your acid pools by 20%.

Level 1

Bragging Rights

Weapon

Hitting enemies with Bladed Chakrams Heals you for 75.

Level 5

Crowd-Pleaser

Combat Trance

Generate 3 Ammo after activating Combat Trance.

Level 3

Instrument of Fate

Heavy Blade

Reduce the Cooldown of Combat Trance by 0.8s for each enemy hit by Heavy Blade.

Level 2

World-traveler

Crouching Tigron

Reset the Cooldown of Crouching Tigron after falling to or below 30% Health.

Level 2

Flying Chakrams

Weapon

Increase your maximum Ammo by 3.

Level 3

Predatory Instincts

Weapon

Bladed Chakrams are able to bounce 2 additional times before disappearing.

Level 1

Antediluvian

Overcharge

Increase the duration of Overcharge by 0.4s.

Level 2

Transient

Rune of Travel

Reduce the cooldown of Rune of Travel by 3s.

Level 3

Primeval Might

Overcharge

Gain 20% Lifesteal while Overcharge is active.

Level 4

Last Remnant

Weapon

Increase your maximum Ammo by 8.

Level 2

Dark Vision

Blitz Upper

Generate 12% Ultimate charge after hitting an enemy with Blitz Upper.

Level 4

Nothing Personal

True Power

Increase your Ultimate charge rate by 15% and enemies at or below 65% Health are Revealed to you.

Level 1

Antediluvian

Overcharge

Increase the duration of Overcharge by 0.4s.

Level 2

Transient

Rune of Travel

Reduce the cooldown of Rune of Travel by 3s.

Level 3

Primeval Might

Overcharge

Gain 20% Lifesteal while Overcharge is active.

Level 4

Last Remnant

Weapon

Increase your maximum Ammo by 8.

Level 2

Dark Vision

Blitz Upper

Generate 12% Ultimate charge after hitting an enemy with Blitz Upper.

Level 4

Nothing Personal

True Power

Increase your Ultimate charge rate by 15% and enemies at or below 65% Health are Revealed to you.

Level 1

Follow the Scent

Fire Spit

Increase the size of your Fire Spit by 48%.

Level 2

Rain of Fire

Thrust

Consume 30% less Booster fuel for 5s after using Thrust.

Level 2

Condescension

Fire Spit

Reduce the Cooldown of Thrust by 2.4s for each enemy hit with Fire Spit.

Level 2

Masterful

Fire Spit

Reduce the Cooldown of Fire Spit by 2.5s for each enemy hit by it.

Level 5

Lung Capacity

Fire Spit

Heal for 200 for each enemy hit with Fire Spit.

Level 4

Combustible

Fire Spit

Fire Spit applies a Knockback and deals an additional 300 damage over 2s.

Level 1

Opulence

Missile Launcher

Heal 240 health for each player hit by Missile Launcher.

Level 4

No Chill

Weapon

Increase your Movement Speed while firing by 10%.

Level 2

Proximity

Advance

Reduce your damage taken by 3% for 3s after activating Advance.

Level 1

Crystal Capacitor

Missile Launcher

Reduce the Cooldown of Missile Launcher by 2.8s.

Level 4

Metal March

Missile Launcher

Reduce your damage taken by 12% for 2s after using Missile Launcher.

Level 4

Rocket Barrage

Missile Launcher

Increase the radius of Missile Launcher's explosions by 66%.

Level 1

A Runaway

Hail of Bombs

Increase the explosion radius of Hail of Bombs' explosives by 30%.

Level 3

Queen's Reign

Cluster Grenade

Reduce the Cooldown of Cluster Grenade by 0.4s for each enemy hit by the initial and secondary explosions.

Level 4

Eureka Moment

Weapon

Generate 1 ammo for every -400 explosive damage done.

Level 2

From Betty with Love

Cluster Grenade

Increase the explosion radius of Cluster Grenade's initial explosion by 12%.

Level 2

Bouncing Betty

Explosive Personality

Heal for 120 for each use of Explosive Personality.

Level 4

Level 1

Favored Quarry

Hunter's Mark

Increase the duration of Hunter's Mark by 1.7999999999999998s.

Level 3

Tracker

Hunter's Mark

Increase your Movement Speed by 20% while Hunter's Mark is active.

Level 4

Ranger

Hunter's Mark

Reduce the Cooldown of Hunter’s Mark by 4s.

Level 4

Primal Might

Armor

Reduce your active Cooldowns by 10% after getting an Elimination.

Level 1

Hunting Season

Hunter's Mark

Heal for 600 after getting a Killing Blow on an enemy affected by Hunter’s Mark.

Level 3

Hunting Party

Hunter's Mark

Hunter's Mark has 2 charges and grants its benefits to allies.

Level 1

Dark Sight

Rend Soul

Every Soul Charge you detonate generates 2 Ammo.

Level 2

Bane

Rend Soul

Every Soul Orb stack you detonate increases your Movement Speed by 3% for 4s, up to a maximum of 15%.

Level 3

Wickedness

Rend Soul

Reduce the Cooldown of Rend Soul by 1.0499999999999998s.

Level 3

Umbral Gait

Weapon

Your Movement Speed is increased by 24% while any enemy is marked with a Soul Orb stack.

Level 3

Sorceress

Weapon

Increase your Ultimate charge rate by 28%.

Level 4

Agony

Restore Soul

For each stack of Soul Orb detonated by Rend Soul, your next Restore Soul ignores 5% of Antihealing (excluding Healing disable effects) up to 25% (5 stacks) for up to 10s.

Level 1

Never Surrender!

Battle Shout

Reduce the Cooldown of Battle Shout by 1.2s.

Level 2

Platemail

Armor

Increase your maximum Health by 450.

Level 3

Ready for War

Bulwark

Increase the recharge rate of your Bulwark by 24%.

Level 4

Shield Wall

Bulwark

Increase the Health of your Bulwark by 400.

Level 2

Hopeguard

Bulwark

Increase your Healing received by 24% while your Bulwark is active.

Level 4

Lian's Shield

Bulwark

Your Shield now regenerates at 120% effectiveness, and regenerates even while it is active.

Level 1

Astral Cycle

Astral Mark

Increase the duration of Astral Mark by 1.6s.

Level 4

Retrograde

Astral Mark

Reduce the Cooldown of Astral Mark by 2s.

Level 5

Relativity

Astral Mark

Heal for 120 every 1s for 10s after applying Astral Mark.

Level 3

Lightyears

Astral Mark

Increase the range of Astral Mark by 8%.

Level 2

Cosmic Barrier

Stellar Wind

Reduce your damage taken by 5% while Stellar Wind is active.

Level 1

Luminary

Astral Mark

All allies already affected by Astral Mark are Healed for an additional 285 when you use Astral Mark.

Level 1