Manifest Destiny

Grace

Increase the travel distance of Grace by 10 units.

Level 2

Swift Jade

Valor

Reduce the Cooldown of Grace by 2.5s for each enemy hit with Valor. Can only happen once every 3s.

Level 5

Inheritance

Armor

Reduce your active Cooldowns by 24% after getting an Elimination.

Level 3

Divine Right

Grace

Gain 20% Movement Speed for 3s after using Grace.

Level 4

Eagle's Emerald

Valor

Generate 1 Ammo for each enemy hit with Valor.

Level 1

Precision

Weapon

Hitting an enemy with Heirloom Rifle increases your Heirloom Rifle damage against that enemy by 7% for 3s, stacking up to 5 times.

Level 1

Healing Vapors

Smoke Screen

Heal for 500 over 2s while in Smoke Screen.

Level 4

Confound

Poison Bolts

Reduce the Cooldown of Smoke Screen by 0.5s for each enemy hit by Poison Bolts.

Level 1

Dissipate

Smoke Screen

Increase your Movement Speed by 40% for 2s after entering Smoke Screen.

Level 5

Poisoner

Poison Bolts

Reduce the Cooldown of Poison Bolts by 3.2s.

Level 4

Quick Smoker

Smoke Screen

Reduce the Cooldown of Smoke Screen by 0.6s.

Level 1

Debilitate

Poison Bolts

Increase the damage done by Poison Bolts by 25% and the rate at which they deal damage by 75%.

Level 1

Masterful

Fire Spit

Reduce the Cooldown of Fire Spit by 2.5s for each enemy hit by it.

Level 5

Condescension

Fire Spit

Reduce the Cooldown of Thrust by 1.2s for each enemy hit with Fire Spit.

Level 1

Survival

Thrust

Reduce the Cooldown of Thrust by 100% after dropping to or below 30% Health.

Level 3

Apex Predator

Armor

Increase your maximum Health by 250.

Level 5

Follow the Scent

Fire Spit

Increase the size of your Fire Spit by 24%.

Level 1

Combustible

Fire Spit

Fire Spit applies a Knockback and deals an additional 300 damage over 2s.

Level 1

Manifest Destiny

Grace

Increase the travel distance of Grace by 10 units.

Level 2

Swift Jade

Valor

Reduce the Cooldown of Grace by 2.5s for each enemy hit with Valor. Can only happen once every 3s.

Level 5

Inheritance

Armor

Reduce your active Cooldowns by 24% after getting an Elimination.

Level 3

Divine Right

Grace

Gain 20% Movement Speed for 3s after using Grace.

Level 4

Eagle's Emerald

Valor

Generate 1 Ammo for each enemy hit with Valor.

Level 1

Precision

Weapon

Hitting an enemy with Heirloom Rifle increases your Heirloom Rifle damage against that enemy by 7% for 3s, stacking up to 5 times.

Level 1

Masterful

Fire Spit

Reduce the Cooldown of Fire Spit by 2.5s for each enemy hit by it.

Level 5

Condescension

Fire Spit

Reduce the Cooldown of Thrust by 1.2s for each enemy hit with Fire Spit.

Level 1

Survival

Thrust

Reduce the Cooldown of Thrust by 100% after dropping to or below 30% Health.

Level 3

Apex Predator

Armor

Increase your maximum Health by 250.

Level 5

Follow the Scent

Fire Spit

Increase the size of your Fire Spit by 24%.

Level 1

Combustible

Fire Spit

Fire Spit applies a Knockback and deals an additional 300 damage over 2s.

Level 1

Moonwalk

Armor

Increase your maximum Moonlight capacity by 40%.

Level 5

Protectors

Guardian Spirit

Regenerate 6% Moonlight every 1s while within 30 units of Luna and not using Moonlight.

Level 3

Moonlight Garden

Weapon

Generate 2% Moonlight for each enemy hit with Light Bow.

Level 1

Restored Faith

Moonlight

Heal for 20 every time you Heal an ally with Moonlight.

Level 2

Sanctum of Faith

Moonlight

Regenerate 4% Moonlight every 1s while not using Moonlight.

Level 4

Sacrifice

Guardian Spirit

[Guardian Spirit] If Io falls below 200 Health while Luna is active, she trades places with Luna, Heals for 50% of her Health, and Luna dies in her stead.

Level 1

Primeval Might

Overcharge

Gain 20% Lifesteal while Overcharge is active.

Level 4

Transient

Rune of Travel

Reduce the cooldown of Rune of Travel by 1s.

Level 1

Spirited

Armor

Increase your maximum Health by 100

Level 2

Evanescent

Rune of Travel

Increase your Movement Speed by 40% while Rune of Travel is active.

Level 5

Guts

Armor

Reduce your damage taken by 15% while at or below 40% Health.

Level 3

Inner Strength

Rune of Travel

Reduce the Cooldown of Overcharge and Blitz Upper by 6s when you Teleport using Rune of Travel.

Level 1

One Step Ahead

Armor

Increase your Movement Speed by 20%.

Level 5

Scapegoat

Deflector Shield

While Deflector Shield is active 8% of damage dealt to you is instead dealt to the Shield.

Level 1

Eyes on the Prize

Precision Sights

Reduce your weapon recoil by 40% while using Precision Sights.

Level 2

Unchecked Ambition

Weapon

Hitting an enemy with a weapon shot has a 60% chance to not consume Ammo.

Level 5

Contingency

Deflector Shield

Increase Deflector Shield's Health by 250.

Level 2

Opportunity in Chaos

Weapon

After firing continuously for 2s, gain 10% increased damage.

Level 1

Level 3

Level 1

Level 2

Level 5

Level 4

Level 1

Flak Jacket

Armor

Increase your maximum Health by 200.

Level 4

Compensator

Iron Sights

Reduce your weapon recoil by 20% while using Iron Sights.

Level 1

Firing Stance

Iron Sights

Increase your Movement Speed by 40% while using Iron Sights.

Level 4

Predator

Iron Sights

Gain 25% Lifesteal while using Iron Sights.

Level 5

Grenadier

Frag Grenade

Reduce the Cooldown of Frag Grenade by 1s for each enemy hit by it.

Level 1

Burst Mode

Weapon

Switch your rifle to Three-Shot-Burst mode, increasing your effective range and firing three 210-damage shots every 0.4s.

Level 1

Level 3

Level 1

Level 2

Level 5

Level 4

Level 1

Manifest Destiny

Grace

Increase the travel distance of Grace by 10 units.

Level 2

Swift Jade

Valor

Reduce the Cooldown of Grace by 2.5s for each enemy hit with Valor. Can only happen once every 3s.

Level 5

Inheritance

Armor

Reduce your active Cooldowns by 24% after getting an Elimination.

Level 3

Divine Right

Grace

Gain 20% Movement Speed for 3s after using Grace.

Level 4

Eagle's Emerald

Valor

Generate 1 Ammo for each enemy hit with Valor.

Level 1

Precision

Weapon

Hitting an enemy with Heirloom Rifle increases your Heirloom Rifle damage against that enemy by 7% for 3s, stacking up to 5 times.

Level 1

Compensator

Iron Sights

Reduce your weapon recoil by 60% while using Iron Sights.

Level 3

Predator

Iron Sights

Gain 5% Lifesteal while using Iron Sights.

Level 1

Flak Jacket

Armor

Increase your maximum Health by 100.

Level 2

XL Mag

Weapon

Increase your maximum Ammo by 10.

Level 5

Firing Stance

Iron Sights

Increase your Movement Speed by 40% while using Iron Sights.

Level 4

Cardio

Hustle

Heal for 300 per second while using Hustle.

Level 1

Level 3

Level 1

Level 2

Level 5

Level 4

Level 1

Level 3

Level 1

Level 2

Level 5

Level 4

Level 1

Depths of Despair

Reduce the damage you take by 12% for 2s after activating Conviction.

Level 3

Tools of Salvation

Increase your maximum Health by 300.

Level 2

Ad Eternum

Reduce the damage you take by 6% while at least one wall segment of Sanctuary is active.

Level 2

Grim Deliverance

Heal for 25 for every percentage point of Ire consumed by abilities.

Level 5

Forged in Battle

Reduce the Cooldown of Sanctuary by 1.7999999999999998s.

Level 3

Persistence

Gain 30% Lifesteal on your hits with Judgement.

Level 1

Depths of Despair

Reduce the damage you take by 12% for 2s after activating Conviction.

Level 3

Tools of Salvation

Increase your maximum Health by 300.

Level 2

Ad Eternum

Reduce the damage you take by 6% while at least one wall segment of Sanctuary is active.

Level 2

Grim Deliverance

Heal for 25 for every percentage point of Ire consumed by abilities.

Level 5

Forged in Battle

Reduce the Cooldown of Sanctuary by 1.7999999999999998s.

Level 3

Persistence

Gain 30% Lifesteal on your hits with Judgement.

Level 1