True Grit

Sniper Mode

Hitting an enemy with a fully-charged Sniper Mode shot Heals you for 200.

Level 4

Beam Me Up

Transporter

Increase the Flight Speed of Transporter by 40%.

Level 4

Prodigy

Sniper Mode

Increase your Movement Speed while in Sniper Mode by 75%.

Level 3

Open Season

Sniper Mode

Hitting an enemy with a fully-charged Sniper Mode shot generates 1 Ammo.

Level 1

Quick Scope

Sniper Mode

Reduce scope-in and scope-out time by 60%.

Level 3

Steady Aim

Sniper Mode

For 4s after hitting a fully-charged weapon shot, your next fully-charged shot will deal an additional 20% damage.

Level 1

Hull Piercer

Harpoon

Reduce the Cooldown of Harpoon by 6s.

Level 5

Sea Dogs

Armor

Increase the Healing you receive by 5% while at or below 50% Health.

Level 1

Hangman's Ire

Armor

Gain a 150-Health Shield after dropping to or below 50% Health.

Level 1

Blow the Man Down

Harpoon

Enemies hit by Harpoon’s Slow have their Movement Speed reduced by an additional 18%.

Level 3

Mortal Skewer

Harpoon

Heal for 350 for each enemy hit with Harpoon.

Level 5

Hurl

Harpoon

Harpoon no longer has a Cooldown, but no longer Slows enemies hit.

Level 1

Arc Lightning

Shock Pulse

Increase the Chain range of Shock Pulse by 40%.

Level 5

Shamanic Might

Armor

Increase your maximum Health by 50.

Level 1

Conduit

Shock Pulse

Increase your Movement Speed by 6% for 3s after hitting an enemy with Shock Pulse.

Level 1

Lightning Rod

Shock Pulse

Heal for 90 for each enemy hit with Shock Pulse.

Level 3

Thunderstruck

Shock Pulse

Generate 30% of your Ammo for each unique enemy hit by Shock Pulse.

Level 5

Maelstrom

Shock Pulse

Increase the damage of Lightning Staff to 70, your weapon shots reduce the Cooldown of Shock Pulse by 0.5s, and Shock Pulse chains 4 additional times. Surgery (+10% Damage)

Level 1

Relentless

Weapon

Reduce the Movement Speed penalty of Scope by 100%.

Level 5

Roost

Scope

Hits with Talon Rifle generate 1% Ultimate charge.

Level 1

Infused Crystals

Weapon

Increase the maximum Ammo count of both your weapons by 3.

Level 3

Grizzled

Armor

Increase your maximum Health by 50.

Level 1

Tactical Retreat

Stealth

Increase your Movement Speed by 50% for 2s after entering Stealth.

Level 5

Nocturnal

Stealth

Reduce the rate of Stealth Energy consumption by 50% and increase your Movement Speed by 20% while in Stealth.

Level 1

Tools of Salvation

Increase your maximum Health by 150.

Level 1

Ad Eternum

Reduce the damage you take by 12% while at least one wall segment of Sanctuary is active.

Level 4

Grim Deliverance

Heal for 20 for every percentage point of Ire consumed by abilities.

Level 4

Eternal Strife

Using an ability above Ire threshold Heals you for 300.

Level 4

Gathering Power

Heal for 140 for each enemy hit with Reckoning.

Level 2

Eternal

Your Ire no longer decays due to not taking or dealing damage, but you no longer gain the passive damage increase while above Ire threshold.

Level 1

Relentless

Weapon

Reduce the Movement Speed penalty of Scope by 100%.

Level 5

Roost

Scope

Hits with Talon Rifle generate 1% Ultimate charge.

Level 1

Infused Crystals

Weapon

Increase the maximum Ammo count of both your weapons by 3.

Level 3

Grizzled

Armor

Increase your maximum Health by 50.

Level 1

Tactical Retreat

Stealth

Increase your Movement Speed by 50% for 2s after entering Stealth.

Level 5

Nocturnal

Stealth

Reduce the rate of Stealth Energy consumption by 50% and increase your Movement Speed by 20% while in Stealth.

Level 1

Astral Cycle

Astral Mark

Increase the duration of Astral Mark by 1.6s.

Level 4

Retrograde

Astral Mark

Reduce the Cooldown of Astral Mark by 2s.

Level 5

Relativity

Astral Mark

Heal for 120 every 1s for 10s after applying Astral Mark.

Level 3

Lightyears

Astral Mark

Increase the range of Astral Mark by 8%.

Level 2

Cosmic Barrier

Stellar Wind

Reduce your damage taken by 5% while Stellar Wind is active.

Level 1

Luminary

Astral Mark

All allies already affected by Astral Mark are Healed for an additional 285 when you use Astral Mark.

Level 1

Relentless

Weapon

Reduce the Movement Speed penalty of Scope by 100%.

Level 5

Roost

Scope

Hits with Talon Rifle generate 1% Ultimate charge.

Level 1

Infused Crystals

Weapon

Increase the maximum Ammo count of both your weapons by 3.

Level 3

Grizzled

Armor

Increase your maximum Health by 50.

Level 1

Tactical Retreat

Stealth

Increase your Movement Speed by 50% for 2s after entering Stealth.

Level 5

Nocturnal

Stealth

Reduce the rate of Stealth Energy consumption by 50% and increase your Movement Speed by 20% while in Stealth.

Level 1

Bloodthirst

Armor

Gain 20% Lifesteal.

Level 5

Excessive Force

Weapon

Increase your maximum Ammo by 15.

Level 5

Ready for War

Bulwark

Increase the recharge rate of your Bulwark by 12%.

Level 2

Vigorous Defense

Bulwark

Heal for 50 every 1s while Bulwark is active.

Level 2

Never Surrender!

Battle Shout

Reduce the Cooldown of Battle Shout by 0.6s.

Level 1

Storm of Bullets

Weapon

Increase your Attack Speed by 40%, but reduce your damage-per-shot by 25%.

Level 1

Otherworldly Tether

Tendril

Gain a 225 Shield for 5s after activating Tendril.

Level 3

The Maw's Embrace

Weapon

Increase your Movement Speed by 8% for 1s after hitting anything with Deadly Scythe.

Level 2

Elegant Paring

Tendril

Reduce the Cooldown of Dark Siphon and Obliteration by 1.6s after activating Tendril.

Level 4

A Change of Faith

Dark Siphon

Reduce the Cooldown of Tendril by 1.2s after hitting an enemy with Dark Siphon.

Level 2

Dance with Fate

Tendril

Reduce the cooldown of Tendril by 2.4s.

Level 4

Relentless Presence

Tendril

Tendril gains a second charge, but its Cooldown is increased by 2s.

Level 1