Antediluvian

Overcharge

Increase the duration of Overcharge by 0.4s.

Level 2

Transient

Rune of Travel

Reduce the cooldown of Rune of Travel by 3s.

Level 3

Primeval Might

Overcharge

Gain 20% Lifesteal while Overcharge is active.

Level 4

Last Remnant

Weapon

Increase your maximum Ammo by 8.

Level 2

Dark Vision

Blitz Upper

Generate 12% Ultimate charge after hitting an enemy with Blitz Upper.

Level 4

Nothing Personal

True Power

Increase your Ultimate charge rate by 15% and enemies at or below 65% Health are Revealed to you.

Level 1

Astral Cycle

Astral Mark

Increase the duration of Astral Mark by 1.6s.

Level 4

Retrograde

Astral Mark

Reduce the Cooldown of Astral Mark by 2s.

Level 5

Relativity

Astral Mark

Heal for 120 every 1s for 10s after applying Astral Mark.

Level 3

Lightyears

Astral Mark

Increase the range of Astral Mark by 8%.

Level 2

Cosmic Barrier

Stellar Wind

Reduce your damage taken by 5% while Stellar Wind is active.

Level 1

Luminary

Astral Mark

All allies already affected by Astral Mark are Healed for an additional 285 when you use Astral Mark.

Level 1

Astral Cycle

Astral Mark

Increase the duration of Astral Mark by 1.6s.

Level 4

Retrograde

Astral Mark

Reduce the Cooldown of Astral Mark by 2s.

Level 5

Relativity

Astral Mark

Heal for 120 every 1s for 10s after applying Astral Mark.

Level 3

Lightyears

Astral Mark

Increase the range of Astral Mark by 8%.

Level 2

Cosmic Barrier

Stellar Wind

Reduce your damage taken by 5% while Stellar Wind is active.

Level 1

Luminary

Astral Mark

All allies already affected by Astral Mark are Healed for an additional 285 when you use Astral Mark.

Level 1

Broken Deity

Guardian Spirit

Increase Luna's deploy range by 20%.

Level 1

Sky-Walker

Armor

Lower your gravity and increase your air control by 30% while firing your Light Bow.

Level 3

Half Moon

Lunar Leap

Reduce the Cooldown of Lunar Leap by 4s.

Level 5

Full Moon

Lunar Leap

Gain a 300-Health Shield for 3s after activating Lunar Leap.

Level 3

Crescent

Lunar Leap

Reset the Cooldown of Lunar Leap after dropping to or below 45% Health.

Level 3

Sacrifice

Guardian Spirit

[Guardian Spirit] If Io falls below 200 Health while Luna is active, she trades places with Luna, Heals for 50% of her Health, and Luna dies in her stead.

Level 1

Fire from Above

Explosive Personality

Increase the vertical movement from each use of Explosive Personality by 40%.

Level 1

Queen's Reign

Cluster Grenade

Reduce the Cooldown of Cluster Grenade by 0.1s for each enemy hit by the initial and secondary explosions.

Level 1

Queen's Entrance

Explosive Personality

Increase the lateral movement from each use of Explosive Personality by 160%.

Level 4

He Will Explode

Hail of Bombs

Reduce the Cooldown of Explosive Personality by 30% after activating Hail of Bombs.

Level 5

Bouncing Betty

Explosive Personality

Heal for 120 for each use of Explosive Personality.

Level 4

Gotta Bounce

Explosive Personality

Reduce the Cooldown of Explosive Personality by 5s, but reduce your Movement Speed by 15%.

Level 1

The Great Worm

Form Swap

Gain a 240-Health Shield for 2s after swapping into Travel Form.

Level 3

Violent Birth

Hardening

Reduce the Cooldown of Acceleration by 1.6s after activating Hardening.

Level 2

Triumphant Return

Form Swap

Reduce your acid consumption by 90% for 0.8s after swapping into Planted Form.

Level 2

Deadly Pursuit

Form Swap

Increase your Movement Speed by 20% for 3s after swapping into Travel Form.

Level 3

Towering Presence

Armor

Increase the Health of your Ultimate Health pool by 1000.

Level 5

Corrosive Acid

Caustic Spray

Augment your acid, increasing the damage done by Caustic Spray to 42, the damage of acid pools to 40, and increase the Movement Speed of allies within your acid pools by 20%.

Level 1

Crow's Nest

Broadside

Hitting yourself with Broadside now applies a Knockback of 300.

Level 2

Expansive Vault

Shortcut

Jump height is increased for your first jump within 1s after exiting a Shortcut.

Level 1

Hangman's Ire

Armor

Gain a 600-Health Shield after dropping to or below 50% Health.

Level 4

Dark Bargain

Shortcut

Heal for 200 every 1s while standing near a Shortcut.

Level 4

Sea Dogs

Armor

Increase the Healing you receive by 20% while at or below 50% Health.

Level 4

Scuttle

Weapon

The projectile launched by reloading your Cursed Howitzer explodes in a 20-unit-radius area on contact.

Level 1

Soul Forge

Restore Soul

Increase the range from the main target at which Restore Soul's secondary Healing can target allies by 18%.

Level 3

Veil

Shadow Travel

Reduce the cast time of Shadow Travel by 30%.

Level 3

Nether Siphon

Shadow Travel

Heal for 150 every 1s while using Shadow Travel.

Level 3

Dusk-Walker

Shadow Travel

Reduce the Cooldown of Shadow Travel by 2.4000000000000004s.

Level 3

Blood Pact

Restore Soul

Heal for 210 every 1s while channeling Restore Soul.

Level 3

Level 1