Level 4

Deadly Pursuit

Form Swap

Increase your Movement Speed by 20% for 2s after swapping into Travel Form.

Level 2

Ancient Wounds

Piercing Quills

Generate 8% of your maximum acid after hitting an enemy with Piercing Quills.

Level 4

Last Efforts

Piercing Quills

Hitting an enemy with Piercing Quills grants 20% Lifesteal for 0.6s.

Level 1

Raze and Burn

Piercing Quills

Heal for 200 after hitting an enemy with Piercing Quills.

Level 4

Unnatural Persistence

Armor

Heal for 25 every 0.25s while on an objective and for 1s after leaving the objective. This effect increases the longer you maintain it, up to 65 every 0.25s.

Level 1

Subjugation

Juggernaut

Increase the distance traveled by Juggernaut by 32%.

Level 4

Shattered Essence

Soul Harvest

Each Soul Fragment gathered Heals you for 15.

Level 1

Harbinger

Soul Harvest

Reduce the Cooldown of Juggernaut by 2s after activating Soul Harvest.

Level 2

Apocalypse

Juggernaut

Heal for 450 for each champion hit by Juggernaut.

Level 3

Declaration of War

Juggernaut

Increase the potency of Juggernaut's Knockup by 35%.

Level 5

Subservience

Soul Harvest

All your living allies are Healed for 1.5% of their maximum Health for each Soul Harvested.

Level 1

Subjugation

Juggernaut

Increase the distance traveled by Juggernaut by 40%.

Level 5

Harbinger

Soul Harvest

Reduce the Cooldown of Juggernaut by 4s after activating Soul Harvest.

Level 4

Void Lord

Soul Harvest

Reduce your damage taken by 5% for 3s after activating Soul Harvest.

Level 1

Shattered Essence

Soul Harvest

Each Soul Fragment gathered Heals you for 15.

Level 1

Apocalypse

Juggernaut

Heal for 600 for each champion hit by Juggernaut.

Level 4

Subservience

Soul Harvest

All your living allies are Healed for 1.5% of their maximum Health for each Soul Harvested.

Level 1

Level 4

Deadly Pursuit

Form Swap

Increase your Movement Speed by 20% for 2s after swapping into Travel Form.

Level 2

Ancient Wounds

Piercing Quills

Generate 8% of your maximum acid after hitting an enemy with Piercing Quills.

Level 4

Last Efforts

Piercing Quills

Hitting an enemy with Piercing Quills grants 20% Lifesteal for 0.6s.

Level 1

Raze and Burn

Piercing Quills

Heal for 200 after hitting an enemy with Piercing Quills.

Level 4

Corrosive Acid

Caustic Spray

Augment your acid, increasing the damage done by Caustic Spray to 42, the damage of acid pools to 40, and increase the Movement Speed of allies within your acid pools by 20%.

Level 1

Collateral Damage

Form Swap

Gain 40% Lifesteal with Caustic Spray for 3s after swapping into Planted Form.

Level 4

Raze and Burn

Piercing Quills

Heal for 200 after hitting an enemy with Piercing Quills.

Level 4

Last Efforts

Piercing Quills

Hitting an enemy with Piercing Quills grants 20% Lifesteal for 0.6s.

Level 1

The Great Worm

Form Swap

Gain a 400-Health Shield for 2s after swapping into Travel Form.

Level 5

Deadly Pursuit

Form Swap

Increase your Movement Speed by 20% for 1s after swapping into Travel Form.

Level 1

Corrosive Acid

Caustic Spray

Augment your acid, increasing the damage done by Caustic Spray to 42, the damage of acid pools to 40, and increase the Movement Speed of allies within your acid pools by 20%.

Level 1

Moonwalk

Armor

Increase your maximum Moonlight capacity by 32%.

Level 4

Restored Faith

Moonlight

Heal for 20 every time you Heal an ally with Moonlight.

Level 2

Spirit Arrows

Moonlight

Reduce the Cooldown of Luna's redeploy by 0.2s after healing with Moonlight.

Level 2

Sky-Walker

Armor

Lower your gravity and increase your air control by 50% while firing your Light Bow.

Level 5

Sanctum of Faith

Moonlight

Regenerate 2% Moonlight every 1s while not using Moonlight.

Level 2

Sacrifice

Guardian Spirit

[Guardian Spirit] If Io falls below 200 Health while Luna is active, she trades places with Luna, Heals for 50% of her Health, and Luna dies in her stead.

Level 1

Extermination

Armor

Reduce the duration and effectiveness of Crowd Control by 25%.

Level 5

Level 2

Towering Presence

Armor

Increase the Health of your Ultimate Health pool by 600.

Level 3

Raze and Burn

Piercing Quills

Heal for 200 after hitting an enemy with Piercing Quills.

Level 4

Last Efforts

Piercing Quills

Hitting an enemy with Piercing Quills grants 20% Lifesteal for 0.6s.

Level 1

Unnatural Persistence

Armor

Heal for 25 every 0.25s while on an objective and for 1s after leaving the objective. This effect increases the longer you maintain it, up to 65 every 0.25s.

Level 1

Void Lord

Soul Harvest

Reduce your damage taken by 15% for 3s after activating Soul Harvest.

Level 3

Sinister Allies

Armor

Increase your maximum Health by 750.

Level 5

Abhorrent Vista

Ignition

Reduce the Cooldown of Ignition by 1.2s.

Level 2

Hellish Lodestones

Ignition

Reduce your damage taken by 12% for 3s after activating Ignition.

Level 4

Tormented Fissure

Ignition

Generate 25 Ammo after activating Ignition.

Level 1

Subservience

Soul Harvest

All your living allies are Healed for 1.5% of their maximum Health for each Soul Harvested.

Level 1

Last Efforts

Piercing Quills

Hitting an enemy with Piercing Quills grants 20% Lifesteal for 0.6s.

Level 1

Level 3

Dreadful Compact

Piercing Quills

Reduce the Cooldown of Hardening by 0.8999999999999999s after hitting an enemy with Piercing Quills.

Level 3

Out of Time

Hardening

Reduce the additional Damage Reduction given by Hardening by half. For 2s after using Hardening, gain its benefits.

Level 4

End of an Era

Hardening

Increase the additional Damage Reduction benefit of Hardening by 80% but reduce its duration by {scale=10|10}%.

Level 4

Unnatural Persistence

Armor

Heal for 25 every 0.25s while on an objective and for 1s after leaving the objective. This effect increases the longer you maintain it, up to 65 every 0.25s.

Level 1

Level 4

Deadly Pursuit

Form Swap

Increase your Movement Speed by 20% for 2s after swapping into Travel Form.

Level 2

Ancient Wounds

Piercing Quills

Generate 8% of your maximum acid after hitting an enemy with Piercing Quills.

Level 4

Last Efforts

Piercing Quills

Hitting an enemy with Piercing Quills grants 20% Lifesteal for 0.6s.

Level 1

Raze and Burn

Piercing Quills

Heal for 200 after hitting an enemy with Piercing Quills.

Level 4

Corrosive Acid

Caustic Spray

Augment your acid, increasing the damage done by Caustic Spray to 42, the damage of acid pools to 40, and increase the Movement Speed of allies within your acid pools by 20%.

Level 1

False Idol

Armor

Increase your Movement Speed by 25% and gain {scale=5|5}% Lifesteal for 2s after falling to or below 40% Health.

Level 5

The Maw's Embrace

Weapon

Increase your Movement Speed by 12% for 1s after hitting anything with Deadly Scythe.

Level 3

What May Come

Tendril

Reduce the damage you take by 8% for 3s after activating Tendril.

Level 2

New Purpose

Armor

Increase your maximum Health by 200.

Level 4

Broken Promises

Obliteration

Heal for 60 for each enemy hit with Obliteration.

Level 1

Level 1

Blood Cannon

Armor

Increase extra Blood Health by 800 while decreasing base Health by {scale=125|125}.

Level 4

Gloomy Nook

Armor

Gain a 150 Shield for 1.75s after falling to or below 75% Health. Can only happen once every 35s.

Level 1

Virulent

Blood Hex

Reduce the Blood cost of Death Wings by 4% while an enemy is marked by Blood Hex.

Level 1

Sharpened Slaughter

Weapon

Increase Projectile Speed by 40%, but set your maximum Ammo to 2.

Level 4

Tainted Form

Weapon

Gain 5% of Max Health when hitting a marked enemy.

Level 5

Level 1

Dreadful Compact

Piercing Quills

Reduce the Cooldown of Hardening by 1.2s after hitting an enemy with Piercing Quills.

Level 4

End of an Era

Hardening

Increase the additional Damage Reduction benefit of Hardening by 60% but reduce its duration by {scale=10|10}%.

Level 3

Raze and Burn

Piercing Quills

Heal for 150 after hitting an enemy with Piercing Quills.

Level 3

Last Efforts

Piercing Quills

Hitting an enemy with Piercing Quills grants 20% Lifesteal for 0.6s.

Level 1

Level 4

Corrosive Acid

Caustic Spray

Augment your acid, increasing the damage done by Caustic Spray to 42, the damage of acid pools to 40, and increase the Movement Speed of allies within your acid pools by 20%.

Level 1