Thrive

Shoulder Bash

Heal for 1000 after hitting an enemy with Shoulder Bash.

Level 5

Watchtower

Siege Shield

Increase the Health of Siege Shield by 250.

Level 1

Trebuchet

Siege Shield

Generate 2 Ammo after activating Siege Shield.

Level 1

Percussion

Kinetic Burst

Increase the Knockback distance of Kinetic Burst by 24%.

Level 3

Indomitable

Armor

Gain 50% Lifesteal.

Level 5

Battering Ram

Shoulder Bash

Reduce your damage taken by 75% while using Shoulder Bash.

Level 1

Raze and Burn

Piercing Quills

Heal for 250 after hitting an enemy with Piercing Quills.

Level 5

Deadly Pursuit

Form Swap

Increase your Movement Speed by 20% for 1s after swapping into Travel Form.

Level 1

Level 2

Out of Time

Hardening

Reduce the additional Damage Reduction given by Hardening by half. For 1.5s after using Hardening, gain its benefits.

Level 3

Last Efforts

Piercing Quills

Hitting an enemy with Piercing Quills grants 20% Lifesteal for 2.4s.

Level 4

Corrosive Acid

Caustic Spray

Augment your acid, increasing the damage done by Caustic Spray to 42, the damage of acid pools to 40, and increase the Movement Speed of allies within your acid pools by 20%.

Level 1

Raze and Burn

Piercing Quills

Heal for 250 after hitting an enemy with Piercing Quills.

Level 5

Deadly Pursuit

Form Swap

Increase your Movement Speed by 20% for 1s after swapping into Travel Form.

Level 1

Level 2

Out of Time

Hardening

Reduce the additional Damage Reduction given by Hardening by half. For 1.5s after using Hardening, gain its benefits.

Level 3

Last Efforts

Piercing Quills

Hitting an enemy with Piercing Quills grants 20% Lifesteal for 2.4s.

Level 4

Corrosive Acid

Caustic Spray

Augment your acid, increasing the damage done by Caustic Spray to 42, the damage of acid pools to 40, and increase the Movement Speed of allies within your acid pools by 20%.

Level 1

Incinerate

Fireball

Reduce the Cooldown of Fireball by 2.25s.

Level 5

Last Stand

Shield

Heal for 90 every 1s while Shield is active and you are at or below 35% Health.

Level 1

Unstoppable Force

Charge

Apply a Knockback of 600 to enemies hit by Charge.

Level 3

Immovable Object

Armor

Increase your Healing received from others by 10% while at or below 50% Health.

Level 2

Brand

Fireball

Heal for 400 for each enemy hit with Fireball.

Level 4

Scorch

Fireball

Increase the damage of Fireball by 35%, decrease its Cooldown by 1s, and deal 20% increased damage for each subsequent target hit after the first.

Level 1

Level 3

Level 2

Level 3

Level 5

Level 2

Level 1

Incinerate

Fireball

Reduce the Cooldown of Fireball by 2.25s.

Level 5

Last Stand

Shield

Heal for 90 every 1s while Shield is active and you are at or below 35% Health.

Level 1

Unstoppable Force

Charge

Apply a Knockback of 600 to enemies hit by Charge.

Level 3

Immovable Object

Armor

Increase your Healing received from others by 10% while at or below 50% Health.

Level 2

Brand

Fireball

Heal for 400 for each enemy hit with Fireball.

Level 4

Scorch

Fireball

Increase the damage of Fireball by 35%, decrease its Cooldown by 1s, and deal 20% increased damage for each subsequent target hit after the first.

Level 1

Thrive

Shoulder Bash

Heal for 1000 after hitting an enemy with Shoulder Bash.

Level 5

Watchtower

Siege Shield

Increase the Health of Siege Shield by 250.

Level 1

Trebuchet

Siege Shield

Generate 2 Ammo after activating Siege Shield.

Level 1

Percussion

Kinetic Burst

Increase the Knockback distance of Kinetic Burst by 24%.

Level 3

Indomitable

Armor

Gain 50% Lifesteal.

Level 5

Battering Ram

Shoulder Bash

Reduce your damage taken by 75% while using Shoulder Bash.

Level 1

Megaton

Blast Shot

Increase the radius of Blast Shot's explosion by 25%.

Level 5

Blast Shower

Blast Shot

Reduce the Cooldown of Blast Shot by 0.5s for each enemy it hits.

Level 1

Incitement

Dodge Roll

Reduce the Cooldown of Dodge Roll by 2s if you hit an enemy with your first weapon shot after using Dodge Roll.

Level 2

Somersault

Armor

Reset the Cooldown of Dodge Roll after dropping to or below 30% Health.

Level 2

Raze

Disengage

Increase the Knockback applied to enemies by Disengage by 40%.

Level 5

Impulse

Blast Shot

Blast Shot has its damage increased by 150 and its Cooldown decreased by 2s.

Level 1

Jolt

Poppy Bomb

Increase the effect of Poppy Bomb’s Knockback by 75% and cause Poppy Bomb to explode on contact with any surface.

Level 5

King's Court

Detonate

Heal for 30 for each Sticky Bomb you Detonate.

Level 2

King's New Cloak

Armor

Increase your maximum Health by 100.

Level 2

Shock and Awe

Detonate

Increase your Movement Speed by 5.5% for 3s after hitting at least one enemy with Detonate.

Level 2

Fallout

Grumpy Bomb

Increase Grumpy Bomb’s Stun duration by 0.4s.

Level 4

Chain Reaction

Detonate

Hitting with more than one Sticky Bomb on a use of Detonate increases the damage of each explosion after the first by 30% of its base damage, up to 150%.

Level 1

Open Season

Sniper Mode

Hitting an enemy with a fully-charged Sniper Mode shot generates 3 Ammo.

Level 3

Octoppressor

Oppressor Mine

Increase the number of targets affected by Oppressor Mines by 4.

Level 4

Amplitude

Oppressor Mine

Increase the range of Oppressor Mine beams by 60%.

Level 3

Beam Me Up

Transporter

Increase the Flight Speed of Transporter by 20%.

Level 2

Calibrate

Weapon

Hitting an enemy with your Sniper Rifle in Carbine mode lowers the Cooldown of Transporter by 0.44999999999999996s.

Level 3

Level 1

Fuel Tank

Armor

Increase your Booster fuel capacity by 30%.

Level 3

Apex Predator

Armor

Increase your maximum Health by 150.

Level 3

Follow the Scent

Fire Spit

Increase the size of your Fire Spit by 96%.

Level 4

Decimate

Salvo

Increase the explosion radius of Salvo by 45%.

Level 3

Propel

Thrust

Restore 20% of your maximum Booster fuel after using Thrust.

Level 2

Fusillade

Weapon

Damage dealt when hitting an enemy directly with Rocket Launcher or Salvo is increased by 25%.

Level 1

Inevitability

Obliteration

Reduce the Cooldown of Dark Siphon by 1.6s after hitting at least one enemy with Obliteration.

Level 4

Elegant Paring

Tendril

Reduce the Cooldown of Dark Siphon and Obliteration by 1.2000000000000002s after activating Tendril.

Level 3

Broken Promises

Obliteration

Heal for 240 for each enemy hit with Obliteration.

Level 4

The Maw's Embrace

Weapon

Increase your Movement Speed by 4% for 1s after hitting anything with Deadly Scythe.

Level 1

Sharpened Resolve

Weapon

Consuming Darkness stacks Heals you for an additional 30 per stack.

Level 3

Level 1