Protectors

Guardian Spirit

Regenerate 2% Moonlight every 1s while within 30 units of Luna and not using Moonlight.

Level 1

Moonwalk

Armor

Increase your maximum Moonlight capacity by 24%.

Level 3

Feral Strength

Guardian Spirit

Increase Luna's Health by 1125.

Level 5

Restored Faith

Moonlight

Heal for 40 every time you Heal an ally with Moonlight.

Level 4

Crescent

Lunar Leap

Reset the Cooldown of Lunar Leap after dropping to or below 30% Health.

Level 2

Sacrifice

Guardian Spirit

[Guardian Spirit] If Io falls below 200 Health while Luna is active, she trades places with Luna, Heals for 50% of her Health, and Luna dies in her stead.

Level 1

Bully

Net Shot

Gain 36% Lifesteal against targets Slowed by Net Shot.

Level 3

Tangled

Net Shot

Increase the duration of Net Shot's Slow by 0.6s.

Level 2

Leg Day

Heroic Leap

Increase the jump strength of Heroic Leap by 10%.

Level 2

Gotcha

Net Shot

Increase your Movement Speed by 40% for 3s after using Net Shot.

Level 4

Concussion

Net Shot

Reduce the Cooldown of Net Shot by 4s.

Level 4

Ensnare

Net Shot

Increase the damage you deal to targets Slowed by Net Shot by 20% and increases the duration of Net Shot's Slow by 0.5s. The damage increase lasts a minimum of 2s.

Level 1

Level 5

Level 2

Level 3

Level 3

Level 2

Level 1

Doomsday

Grumpy Bomb

Heal for 520 after using Grumpy Bomb.

Level 4

Fallout

Grumpy Bomb

Increase Grumpy Bomb’s Stun duration by 0.5s.

Level 5

Countdown

Grumpy Bomb

Reduce the Cooldown of Grumpy Bomb by 1.6s.

Level 2

Mad Bomber

Grumpy Bomb

Increase your Movement Speed by 14% for 3s after using Grumpy Bomb.

Level 2

Uncontrolled

Armor

Reduce your active Cooldowns by 16% after getting an Elimination.

Level 2

Accelerant

Grumpy Bomb

Reduces the time it takes for Grumpy Bomb to explode by 40%.

Level 1

Electrostatic

Weapon

Increase your Ammo regeneration speed by 36%.

Level 3

Arc Lightning

Shock Pulse

Increase the Chain range of Shock Pulse by 40%.

Level 5

Totemic Rescue

Armor

Gain a 450-Health Shield for 3s after dropping to or below 30% Health.

Level 3

Thunderlord

Weapon

Increase your maximum Ammo by 20%.

Level 2

Thunderstruck

Shock Pulse

Generate 12% of your Ammo for each unique enemy hit by Shock Pulse.

Level 2

Maelstrom

Shock Pulse

Increase the damage of Lightning Staff to 70, your weapon shots reduce the Cooldown of Shock Pulse by 0.5s, and Shock Pulse chains 4 additional times. Surgery (+10% Damage)

Level 1

Overflowing Delights

Swarm

Increase the Radius of Swarm by 6%, but increase its cost by {scale=3|3}%.

Level 3

Tainted Form

Weapon

Gain 2% of Max Health when hitting a marked enemy.

Level 2

Blood Sense

Blood Hex

Hitting an enemy with Blood Hex will Reveal them to your team for 1.5s.

Level 3

Enriched Blood

Swarm

Increase the damage and Healing of Swarm by 12%, but reduce its duration by {scale=0.35|0.35}s.

Level 3

Symbiotic Relationship

Swarm

Increase the Healing allies receive from Swarm by 16%, but increase its cost by {scale=4|4}%.

Level 4

Maelstrom of Carnage

Swarm

Swarm is now cast on yourself and follows you around instead of being placed on the ground, its ability cost is decreased by 25%, and its radius is increased by 30%.

Level 1

Level 5

Level 2

Level 3

Level 3

Level 2

Level 1

Ad Eternum

Reduce the damage you take by 12% while at least one wall segment of Sanctuary is active.

Level 4

Righteous Fury

Reduce the Cooldown of Sanctuary by 1.2s after using Reckoning.

Level 2

Solemn Watch

Increase the distance traveled by Conviction by 12%.

Level 2

Flames of Wrath

Increase the duration of the Lift applied by Reckoning by 20%.

Level 2

Overwhelming Presence

Heal for 60 every 0.5s while at least one wall segment of Sanctuary is active.

Level 5

Persistence

Gain 30% Lifesteal on your hits with Judgement.

Level 1

Grizzled

Armor

Increase your maximum Health by 100.

Level 2

Tactical Retreat

Stealth

Increase your Movement Speed by 30% for 2s after entering Stealth.

Level 3

Escape Plan

Stealth

Heal for 300 after entering Stealth at or below 65% Health.

Level 3

Relentless

Weapon

Reduce the Movement Speed penalty of Scope by 40%.

Level 2

Roost

Scope

Hits with Talon Rifle generate 5% Ultimate charge.

Level 5

Nocturnal

Stealth

Reduce the rate of Stealth Energy consumption by 50% and increase your Movement Speed by 20% while in Stealth.

Level 1

Old Wounds

Armor

Increase your maximum Health by 150.

Level 1

Hell Hunter

Weapon

Generate 1 Ammo after hitting a fully-charged weapon shot.

Level 1

Paradox

Second Chance

Reduce the Cooldown of Second Chance by 2.5s.

Level 5

Beyond The Veil

Stasis Field

Passing through Stasis Field increases your allies' or your own Movement Speed by 28% for 3s.

Level 4

Continuum Shift

Second Chance

Second Chance sends you 1.6s further into the past.

Level 4

Unstable Fissure

Second Chance

When you use Second Chance you also Rewind enemies within 20 units 3s into the past.

Level 1

Level 5

Level 2

Level 3

Level 3

Level 2

Level 1

Protectors

Guardian Spirit

Regenerate 2% Moonlight every 1s while within 30 units of Luna and not using Moonlight.

Level 1

Moonwalk

Armor

Increase your maximum Moonlight capacity by 24%.

Level 3

Feral Strength

Guardian Spirit

Increase Luna's Health by 1125.

Level 5

Restored Faith

Moonlight

Heal for 40 every time you Heal an ally with Moonlight.

Level 4

Crescent

Lunar Leap

Reset the Cooldown of Lunar Leap after dropping to or below 30% Health.

Level 2

Sacrifice

Guardian Spirit

[Guardian Spirit] If Io falls below 200 Health while Luna is active, she trades places with Luna, Heals for 50% of her Health, and Luna dies in her stead.

Level 1

Strongarm

Dredge Anchor

Reduce the Cooldown of Dredge Anchor by 3s.

Level 5

Sea Legs

Dredge Anchor

Heal for 120 after hitting an enemy with Dredge Anchor.

Level 1

Salvage

Dredge Anchor

Regenerate 1 Ammo every 0.5s for 2s after activating Dredge Anchor.

Level 4

Carapace

Shell Shield

Increase the Health of Shell Shield by 600.

Level 3

Tidal Grace

Shell Shield

Heal for 110 every 1s while Shell Shield is active.

Level 2

Half Shell

Shell Shield

Shell Shield is placed on the ground instead of channeled around you, and lasts 2s longer.

Level 1

Level 5

Level 2

Level 3

Level 3

Level 2

Level 1

Level 5

Level 2

Level 3

Level 3

Level 2

Level 1