Reset the Cooldown of Lunar Leap after dropping to or below 30% Health.
Level 2
Feral Strength
Guardian Spirit
Increase Luna's Health by 450.
Level 2
Sky-Walker
Armor
Lower your gravity and increase your air control by 10% while firing your Light Bow.
Level 1
Moonwalk
Armor
Increase your maximum Moonlight capacity by 40%.
Level 5
Restored Faith
Moonlight
Heal for 50 every time you Heal an ally with Moonlight.
Level 5
Sacrifice
Guardian Spirit
[Guardian Spirit] If Io falls below 200 Health while Luna is active, she trades places with Luna, Heals for 50% of her Health, and Luna dies in her stead.