Extermination

Armor

Reduce the duration and effectiveness of Crowd Control by 15%.

Level 3

Deadly Predator

Weapon

Increase your maximum acid capacity by 8%.

Level 1

Raze and Burn

Piercing Quills

Heal for 50 after hitting an enemy with Piercing Quills.

Level 1

The Great Worm

Form Swap

Gain a 400-Health Shield for 2s after swapping into Travel Form.

Level 5

Towering Presence

Armor

Increase the Health of your Ultimate Health pool by 1000.

Level 5

Corrosive Acid

Caustic Spray

Augment your acid, increasing the damage done by Caustic Spray to 42, the damage of acid pools to 40, and increase the Movement Speed of allies within your acid pools by 20%.

Level 1

Tools of Salvation

Increase your maximum Health by 150.

Level 1

Ad Eternum

Reduce the damage you take by 12% while at least one wall segment of Sanctuary is active.

Level 4

Grim Deliverance

Heal for 20 for every percentage point of Ire consumed by abilities.

Level 4

Eternal Strife

Using an ability above Ire threshold Heals you for 300.

Level 4

Gathering Power

Heal for 140 for each enemy hit with Reckoning.

Level 2

Persistence

Gain 30% Lifesteal on your hits with Judgement.

Level 1

Grim Deliverance

Heal for 20 for every percentage point of Ire consumed by abilities.

Level 4

Depths of Despair

Reduce the damage you take by 12% for 2s after activating Conviction.

Level 3

Ad Eternum

Reduce the damage you take by 9% while at least one wall segment of Sanctuary is active.

Level 3

True Conviction

Reduce the damage your other allies take by 10% while they are within 35 units of the middle wall segment of your Sanctuary.

Level 2

Eternal Strife

Using an ability above Ire threshold Heals you for 225.

Level 3

Tempering

Augment Reckoning, Sanctuary, and Conviction, but increase their base Cooldown by 25%.

Level 1

Grim Deliverance

Heal for 20 for every percentage point of Ire consumed by abilities.

Level 4

Depths of Despair

Reduce the damage you take by 12% for 2s after activating Conviction.

Level 3

Ad Eternum

Reduce the damage you take by 9% while at least one wall segment of Sanctuary is active.

Level 3

True Conviction

Reduce the damage your other allies take by 10% while they are within 35 units of the middle wall segment of your Sanctuary.

Level 2

Eternal Strife

Using an ability above Ire threshold Heals you for 225.

Level 3

Tempering

Augment Reckoning, Sanctuary, and Conviction, but increase their base Cooldown by 25%.

Level 1

Run and Gun

Hustle

Regenerate 4 Ammo every 1s while using Hustle.

Level 2

Scramble

Hustle

Reduce damage taken by 12% while using Hustle.

Level 4

Hot Potato

Frag Grenade

Reduce the cook time of Frag Grenade by 20%.

Level 2

Second Wind

Hustle

Increase your Movement Speed after deactivating Hustle by 30% for 1.5s.

Level 3

Guerrilla Warfare

Hustle

Reduce the Cooldown of Frag Grenade by 0.8s every 1s while Hustle is active.

Level 4

Cardio

Hustle

Heal for 300 per second while using Hustle.

Level 1

Salvage

Dredge Anchor

Regenerate 1 Ammo every 0.5s for 0.5s after activating Dredge Anchor.

Level 1

Surf

Shell Spin

Increase the travel distance of Shell Spin by 30%.

Level 3

Ebb and Flow

Armor

Reduce the Cooldown of Shell Spin by 60% after falling to or below 40% Health.

Level 3

Harden

Dredge Anchor

Reduce your damage taken by 15% for 2s after hitting an enemy with Dredge Anchor.

Level 3

Strongarm

Dredge Anchor

Reduce the Cooldown of Dredge Anchor by 3s.

Level 5

Pluck

Dredge Anchor

For 4s after hitting an enemy with Dredge Anchor, your next weapon shot against that enemy deals 75% increased damage.

Level 1