Feral Strength

Guardian Spirit

Increase Luna's Health by 900.

Level 4

Swift Arrows

Moonlight

Increase allies' Movement Speed by 18% while they are being Healed by Moonlight.

Level 3

Restored Faith

Moonlight

Heal for 20 every time you Heal an ally with Moonlight.

Level 2

Celestial Body

Armor

Increase your maximum Health by 150.

Level 3

Full Moon

Lunar Leap

Gain a 300-Health Shield for 3s after activating Lunar Leap.

Level 3

Sacrifice

Guardian Spirit

[Guardian Spirit] If Io falls below 200 Health while Luna is active, she trades places with Luna, Heals for 50% of her Health, and Luna dies in her stead.

Level 1

Inevitability

Obliteration

Reduce the Cooldown of Dark Siphon by 0.8s after hitting at least one enemy with Obliteration.

Level 2

A Broken Path

Dark Siphon

Increase the Healing received from Dark Siphon by 200 health.

Level 4

New Purpose

Armor

Increase your maximum Health by 50.

Level 1

Unified in Purpose

Dark Siphon

Reduce the damage you take by 10% for 3s after hitting an enemy with Dark Siphon.

Level 3

Crimson Ascent

Dark Siphon

Reduce the Cooldown of Dark Siphon by 5s if it fails to hit a target.

Level 5

Deafening Silence

Obliteration

Obliteration now Silences enemies hit for 1.65s.

Level 1

Heavy Metal

Armor

Increase your maximum Health by 300.

Level 2

Battlement

Siege Shield

Increase the duration of Siege Shield by 1.2s.

Level 2

Siege Engine

Kinetic Burst

Reduce the Cooldown of Siege Shield by 5s for each enemy hit with Kinetic Burst.

Level 5

Ramparts

Siege Shield

Reduce the Cooldown of Siege Shield by 1.6s.

Level 2

Indomitable

Armor

Gain 40% Lifesteal.

Level 4

Battering Ram

Shoulder Bash

Reduce your damage taken by 75% while using Shoulder Bash.

Level 1

Inevitability

Obliteration

Reduce the Cooldown of Dark Siphon by 1.6s after hitting at least one enemy with Obliteration.

Level 4

Elegant Paring

Tendril

Reduce the Cooldown of Dark Siphon and Obliteration by 1.2000000000000002s after activating Tendril.

Level 3

Broken Promises

Obliteration

Heal for 240 for each enemy hit with Obliteration.

Level 4

New Purpose

Armor

Increase your maximum Health by 50.

Level 1

Sharpened Resolve

Weapon

Consuming Darkness stacks Heals you for an additional 30 per stack.

Level 3

Unyielding Pressure

Your Dark Siphon now deals an additional 10% of the target's maximum Health as damage over 1.5s.

Level 1

Feral Strength

Guardian Spirit

Increase Luna's Health by 900.

Level 4

Swift Arrows

Moonlight

Increase allies' Movement Speed by 18% while they are being Healed by Moonlight.

Level 3

Restored Faith

Moonlight

Heal for 20 every time you Heal an ally with Moonlight.

Level 2

Celestial Body

Armor

Increase your maximum Health by 150.

Level 3

Full Moon

Lunar Leap

Gain a 300-Health Shield for 3s after activating Lunar Leap.

Level 3

Sacrifice

Guardian Spirit

[Guardian Spirit] If Io falls below 200 Health while Luna is active, she trades places with Luna, Heals for 50% of her Health, and Luna dies in her stead.

Level 1

Heavy Metal

Armor

Increase your maximum Health by 300.

Level 2

Battlement

Siege Shield

Increase the duration of Siege Shield by 1.2s.

Level 2

Siege Engine

Kinetic Burst

Reduce the Cooldown of Siege Shield by 5s for each enemy hit with Kinetic Burst.

Level 5

Ramparts

Siege Shield

Reduce the Cooldown of Siege Shield by 1.6s.

Level 2

Indomitable

Armor

Gain 40% Lifesteal.

Level 4

Battering Ram

Shoulder Bash

Reduce your damage taken by 75% while using Shoulder Bash.

Level 1

Contingency

Deflector Shield

Increase Deflector Shield's Health by 125.

Level 1

One Step Ahead

Armor

Increase your Movement Speed by 20%.

Level 5

Unchecked Ambition

Weapon

Hitting an enemy with a weapon shot has a 48% chance to not consume Ammo.

Level 4

Runic Ammunition

Weapon

Gain 16% Lifesteal.

Level 4

Hidden Reserves

Precision Sights

Every weapon shot made while Precision Sights is active has an 8% chance to not consume Ammo.

Level 1

Suspect Everyone

Armor

The Health of Deflector Shield is increased by 350 and its Cooldown is reduced by half if it is destroyed by an enemy.

Level 1

True Grit

Sniper Mode

Hitting an enemy with a fully-charged Sniper Mode shot Heals you for 200.

Level 4

Beam Me Up

Transporter

Increase the Flight Speed of Transporter by 40%.

Level 4

Prodigy

Sniper Mode

Increase your Movement Speed while in Sniper Mode by 75%.

Level 3

Open Season

Sniper Mode

Hitting an enemy with a fully-charged Sniper Mode shot generates 1 Ammo.

Level 1

Quick Scope

Sniper Mode

Reduce scope-in and scope-out time by 60%.

Level 3

Reposition

Transporter

Using Transporter instantly Teleports you to its target location.

Level 1