Lightyears

Astral Mark

Increase the range of Astral Mark by 16%.

Level 4

Penumbra

Weapon

Gain 5% Lifesteal.

Level 1

Relativity

Astral Mark

Heal for 80 every 1s for 10s after applying Astral Mark.

Level 2

Retrograde

Astral Mark

Reduce the Cooldown of Astral Mark by 1.2000000000000002s.

Level 3

Astral Cycle

Astral Mark

Increase the duration of Astral Mark by 2s.

Level 5

Luminary

Astral Mark

All allies already affected by Astral Mark are Healed for an additional 285 when you use Astral Mark.

Level 1

Grim Deliverance

Heal for 25 for every percentage point of Ire consumed by abilities.

Level 5

Tools of Salvation

Increase your maximum Health by 150.

Level 1

Depths of Despair

Reduce the damage you take by 8% for 2s after activating Conviction.

Level 2

Forged in Battle

Reduce the Cooldown of Sanctuary by 1.7999999999999998s.

Level 3

Ad Eternum

Reduce the damage you take by 12% while at least one wall segment of Sanctuary is active.

Level 4

Persistence

Gain 30% Lifesteal on your hits with Judgement.

Level 1

Full Moon

Lunar Leap

Gain a 500-Health Shield for 3s after activating Lunar Leap.

Level 5

Eternal Reliquary

Lunar Leap

Generate 4 Ammo after using Lunar Leap.

Level 2

Half Moon

Lunar Leap

Reduce the Cooldown of Lunar Leap by 0.8s.

Level 1

Celestial Body

Armor

Increase your maximum Health by 200.

Level 4

Crescent

Lunar Leap

Reset the Cooldown of Lunar Leap after dropping to or below 45% Health.

Level 3

Sacrifice

Guardian Spirit

[Guardian Spirit] If Io falls below 200 Health while Luna is active, she trades places with Luna, Heals for 50% of her Health, and Luna dies in her stead.

Level 1

Twilight

Dead Zone

Reduce the Cooldown of Dead Zone by 4s.

Level 5

Sprouts

Seedling

Reduce the Cooldown of Seedling by 1.5s for each enemy hit by it.

Level 3

Spritely

Flutter

Reduce your damage taken by 5% for 3s after activating Flutter.

Level 1

Hijinks

Flutter

Heal for 50 after activating Flutter.

Level 1

Just Believe!

Armor

Reduce your active Cooldowns by 40% after getting an Elimination.

Level 5

Nightshade

Dead Zone

Dead Zone damage is increased to 650.

Level 1

Triumphant Return

Form Swap

Reduce your acid consumption by 90% for 0.4s after swapping into Planted Form.

Level 1

Collateral Damage

Form Swap

Gain 50% Lifesteal with Caustic Spray for 3s after swapping into Planted Form.

Level 5

Ancient Wounds

Piercing Quills

Generate 6% of your maximum acid after hitting an enemy with Piercing Quills.

Level 3

Raze and Burn

Piercing Quills

Heal for 250 after hitting an enemy with Piercing Quills.

Level 5

Deadly Pursuit

Form Swap

Increase your Movement Speed by 20% for 1s after swapping into Travel Form.

Level 1

Sight Begets Strength

Primal Vision

Allied players within range of your Primal Vision gain a stackable 90-Health Shield and 15% increased Movement Speed for 4s each time you pulse, but your Primal Vision has its Cooldown increased by 2s.

Level 1

Moonlight Garden

Weapon

Generate 4% Moonlight for each enemy hit with Light Bow.

Level 2

Moonwalk

Armor

Increase your maximum Moonlight capacity by 40%.

Level 5

Crescent

Lunar Leap

Reset the Cooldown of Lunar Leap after dropping to or below 30% Health.

Level 2

Protectors

Guardian Spirit

Regenerate 10% Moonlight every 1s while within 30 units of Luna and not using Moonlight.

Level 5

Restored Faith

Moonlight

Heal for 10 every time you Heal an ally with Moonlight.

Level 1

Goddess' Blessing

Moonlight

Allies being Healed by Moonlight take 15% reduced damage.

Level 1

In The Fray

Nade Launcher

Decrease your damage taken by 20% for 3s after using Nade Launcher.

Level 5

Turn The Table

Nade Launcher

Heal for 180 for each enemy hit with Nade Launcher.

Level 3

Favored Quarry

Hunter's Mark

Increase the duration of Hunter's Mark by 1.2s.

Level 2

Hunting Season

Hunter's Mark

Heal for 400 after getting a Killing Blow on an enemy affected by Hunter’s Mark.

Level 2

Primal Might

Armor

Reduce your active Cooldowns by 30% after getting an Elimination.

Level 3

Mercy Kill

Nade Launcher

Nade Launcher has 3 charges, but reduce the damage of each to 650.

Level 1

Lightyears

Astral Mark

Increase the range of Astral Mark by 16%.

Level 4

Penumbra

Weapon

Gain 5% Lifesteal.

Level 1

Relativity

Astral Mark

Heal for 80 every 1s for 10s after applying Astral Mark.

Level 2

Retrograde

Astral Mark

Reduce the Cooldown of Astral Mark by 1.2000000000000002s.

Level 3

Astral Cycle

Astral Mark

Increase the duration of Astral Mark by 2s.

Level 5

Luminary

Astral Mark

All allies already affected by Astral Mark are Healed for an additional 285 when you use Astral Mark.

Level 1

Enriched Blood

Swarm

Increase the damage and Healing of Swarm by 20%, but reduce its duration by {scale=0.35|0.35}s.

Level 5

Gloomy Nook

Armor

Gain a 150 Shield for 1.75s after falling to or below 75% Health. Can only happen once every 35s.

Level 1

Unholy Flight

Death Wings

Reduce your damage taken by 5% for 1.5s after activating Death Wings.

Level 1

Symbiotic Relationship

Swarm

Increase the Healing allies receive from Swarm by 20%, but increase its cost by {scale=4|4}%.

Level 5

Waning Moon

Blood Hex

Reduce the Blood cost of Swarm by 15% while an ally is marked by Blood Hex.

Level 3

Cursed Accord

Armor

Decrease the Blood cost of your abilities by 35% and increase your Healing done by 15%, but reduce the damage you do by 25%.

Level 1

Looks That Kill

Fireball

Reduce the Cooldown of Charge by 2.4s after hitting an enemy with Fireball.

Level 4

Heat Transfer

Shield

Reduce the Cooldown of Charge by 1s for every 1000 damage your Shield takes.

Level 2

Last Stand

Shield

Heal for 90 every 1s while Shield is active and you are at or below 35% Health.

Level 1

Cavalier

Armor

Increase your maximum Health by 600.

Level 4

Incinerate

Fireball

Reduce the Cooldown of Fireball by 1.8s.

Level 4

Formidable

Charge

You now have 2 charges of Charge, each Charge deals an additional 100 damage, and you are Immune to Crowd Control while Charging, but the Cooldown of Charge is increased to 13s.

Level 1

Healing Station

Turret

Heal for 75 every 1s while standing near any of your Turrets.

Level 3

Bunker

Barricade

Increase the Health of Barricade by 800.

Level 4

Failsafe

Armor

Reset the Cooldown of Rocket Boots after falling to or below 30% Health.

Level 3

Double-Time

Rocket Boots

Heal for 150 every 1s for 3s after activating Rocket Boots.

Level 3

Bowling Ball

Rocket Boots

Gain a 300-Health Shield for 3s after activating Rocket Boots.

Level 2

Fortify

Barricade

Increase the maximum Health of Barricade by 2000 and reduce its Cooldown by 3s.

Level 1

False Idol

Armor

Increase your Movement Speed by 5% and gain {scale=5|5}% Lifesteal for 2s after falling to or below 40% Health.

Level 1

A Broken Path

Dark Siphon

Increase the Healing received from Dark Siphon by 200 health.

Level 4

Unified in Purpose

Dark Siphon

Reduce the damage you take by 10% for 3s after hitting an enemy with Dark Siphon.

Level 3

Sharpened Resolve

Weapon

Consuming Darkness stacks Heals you for an additional 40 per stack.

Level 4

What May Come

Tendril

Reduce the damage you take by 12% for 3s after activating Tendril.

Level 3

Relentless Presence

Tendril

Tendril gains a second charge, but its Cooldown is increased by 2s.

Level 1

In The Fray

Nade Launcher

Decrease your damage taken by 20% for 3s after using Nade Launcher.

Level 5

Turn The Table

Nade Launcher

Heal for 180 for each enemy hit with Nade Launcher.

Level 3

Favored Quarry

Hunter's Mark

Increase the duration of Hunter's Mark by 1.2s.

Level 2

Hunting Season

Hunter's Mark

Heal for 400 after getting a Killing Blow on an enemy affected by Hunter’s Mark.

Level 2

Primal Might

Armor

Reduce your active Cooldowns by 30% after getting an Elimination.

Level 3

Mercy Kill

Nade Launcher

Nade Launcher has 3 charges, but reduce the damage of each to 650.

Level 1

Standing Stones

Warder's Field

Reduce the Cooldown of Warder's Field by 3s.

Level 3

Stone Bulwark

Earthen Guard

Heal for 28 every 1s while Earthen Guard is active.

Level 1

Caretaker

Warder's Field

Heal for 110 every 1s while standing in Warder's Field.

Level 5

Insurmountable

Armor

Increase your maximum Ammo by 2.

Level 2

Sacred Ground

Warder's Field

Reduce your damage taken by 20% while standing in Warder's Field.

Level 4

Treacherous Ground

Warder's Field

The radius of Warder's Field is increased by 50% and it Cripples enemies within it.

Level 1

Viciousness

Counter

Reduce the Cooldown of Counter by 75% after successfully hitting an enemy with Counter.

Level 5

Master Stance

Counter

Reduce the Cooldown of Whirl by 2s after successfully hitting an enemy with Counter.

Level 2

Embers

Billow

Regenerate 2 Ammo every 0.5s for 1s after activating Billow.

Level 2

Fan the Flames

Billow

Reduce the Cooldown of Billow by 28% after getting an Elimination.

Level 2

Perfect Block

Counter

Heal for 400 after using Counter.

Level 4

Smolder

Billow

Heal for 15% of your maximum Health per second while using Billow.

Level 1

Confound

Poison Bolts

Reduce the Cooldown of Smoke Screen by 0.5s for each enemy hit by Poison Bolts.

Level 1

Emergency Exit

Hidden

Automatically enter Stealth after dropping to or below 30% Health. This can happen once every -5s.

Level 1

Healing Vapors

Smoke Screen

Heal for 625 over 2s while in Smoke Screen.

Level 5

Decrepify

Poison Bolts

You do not consume Ammo for 2s after hitting an enemy with Poison Bolts.

Level 4

Shadow Affinity

Hidden

Reduce the Cooldown of Hidden by 4s.

Level 4

Debilitate

Poison Bolts

Increase the damage done by Poison Bolts by 25% and the rate at which they deal damage by 75%.

Level 1