Conduction

Recharge

Reduce the Cooldown of Recharge by 2s.

Level 5

Glyph of Health

Armor

Increase your maximum Health by 150.

Level 1

Arcane Etching

Nullify

Gain a 100-Health Shield after activating Nullify.

Level 1

Timeshaper

Protection

Generate 6 Ammo for the target of your Protection.

Level 3

Hearthwarder

Protection

Generate 375 Shield after using Protection.

Level 5

Direct Current

Nullify

Double the range at which you can use Nullify.

Level 1

Triumphant Return

Form Swap

Reduce your acid consumption by 90% for 0.8s after swapping into Planted Form.

Level 2

Last Efforts

Piercing Quills

Hitting an enemy with Piercing Quills grants 20% Lifesteal for 2.4s.

Level 4

Extermination

Armor

Reduce the duration and effectiveness of Crowd Control by 10%.

Level 2

The Great Worm

Form Swap

Gain a 240-Health Shield for 2s after swapping into Travel Form.

Level 3

Towering Presence

Armor

Increase the Health of your Ultimate Health pool by 800.

Level 4

Sight Begets Strength

Primal Vision

Allied players within range of your Primal Vision gain a stackable 90-Health Shield and 15% increased Movement Speed for 4s each time you pulse, but your Primal Vision has its Cooldown increased by 2s.

Level 1

Conduction

Recharge

Reduce the Cooldown of Recharge by 2s.

Level 5

Glyph of Health

Armor

Increase your maximum Health by 150.

Level 1

Arcane Etching

Nullify

Gain a 100-Health Shield after activating Nullify.

Level 1

Timeshaper

Protection

Generate 6 Ammo for the target of your Protection.

Level 3

Hearthwarder

Protection

Generate 375 Shield after using Protection.

Level 5

Field Study

Protection

Increase the damage done by allies by 10% while they are Shielded by Protection.

Level 1

Grenadier

Frag Grenade

Reduce the Cooldown of Frag Grenade by 5s for each enemy hit by it.

Level 5

Fire in the Hole

Frag Grenade

Generate 12 Ammo for each enemy hit with Frag Grenade.

Level 2

Run and Gun

Hustle

Regenerate 2 Ammo every 1s while using Hustle.

Level 1

Guerrilla Warfare

Hustle

Reduce the Cooldown of Frag Grenade by 1s every 1s while Hustle is active.

Level 5

Predator

Iron Sights

Gain 10% Lifesteal while using Iron Sights.

Level 2

Shrapnel

Frag Grenade

Increase the radius of Frag Grenade's explosion by 40%.

Level 1

Conduction

Recharge

Reduce the Cooldown of Recharge by 2s.

Level 5

Glyph of Health

Armor

Increase your maximum Health by 150.

Level 1

Arcane Etching

Nullify

Gain a 100-Health Shield after activating Nullify.

Level 1

Timeshaper

Protection

Generate 6 Ammo for the target of your Protection.

Level 3

Hearthwarder

Protection

Generate 375 Shield after using Protection.

Level 5

Field Study

Protection

Increase the damage done by allies by 10% while they are Shielded by Protection.

Level 1

Watchtower

Siege Shield

Increase the Health of Siege Shield by 750.

Level 3

Brawl

Kinetic Burst

Heal for 300 over 2s for each enemy hit by Kinetic Burst.

Level 2

Trebuchet

Siege Shield

Generate 4 Ammo after activating Siege Shield.

Level 2

Indomitable

Armor

Gain 40% Lifesteal.

Level 4

Percussion

Kinetic Burst

Increase the Knockback distance of Kinetic Burst by 32%.

Level 4

Fortress Breaker

Siege Shield

Increase Siege Shield's maximum Health by 2500 and its size by 50%, but reduce its Movement Speed by 80%.

Level 1

Grim Deliverance

Heal for 20 for every percentage point of Ire consumed by abilities.

Level 4

Depths of Despair

Reduce the damage you take by 12% for 2s after activating Conviction.

Level 3

Ad Eternum

Reduce the damage you take by 9% while at least one wall segment of Sanctuary is active.

Level 3

Gathering Power

Heal for 140 for each enemy hit with Reckoning.

Level 2

Eternal Strife

Using an ability above Ire threshold Heals you for 225.

Level 3

Tempering

Augment Reckoning, Sanctuary, and Conviction, but increase their base Cooldown by 25%.

Level 1

Retrograde

Astral Mark

Reduce the Cooldown of Astral Mark by 2s.

Level 5

Penumbra

Weapon

Gain 5% Lifesteal.

Level 1

Lightyears

Astral Mark

Increase the range of Astral Mark by 8%.

Level 2

Relativity

Astral Mark

Heal for 120 every 1s for 10s after applying Astral Mark.

Level 3

Astral Cycle

Astral Mark

Increase the duration of Astral Mark by 1.6s.

Level 4

Luminary

Astral Mark

All allies already affected by Astral Mark are Healed for an additional 285 when you use Astral Mark.

Level 1

Frontline Commander

Increase allies and decrease enemies' Movement Speed by 35% for 4 seconds when passing through Abyssal Fortress.

Level 5

Brutal Pursuer

Rift Slash

Increase your Movement Speed by 8% for 5s after casting Rift Slash.

Level 2

Unyielding Advance

Increase Abyssal Fortress' Health by 400

Level 2

Unbreakable Will

Reduce the Health reduction from placing Abyssal Fortress >30 units away by 80%.

Level 4

Forever Changed

Increase your maximum Health by 300.

Level 2

Abyssal Breach

Enemies that pass through Abyssal Fortress are Revealed and take 10% additional weapon damage from you and your allies for 3s. Reduce Abyssal Fortress' Cooldown by 2s.

Level 1

Retrograde

Astral Mark

Reduce the Cooldown of Astral Mark by 2s.

Level 5

Penumbra

Weapon

Gain 5% Lifesteal.

Level 1

Lightyears

Astral Mark

Increase the range of Astral Mark by 8%.

Level 2

Relativity

Astral Mark

Heal for 120 every 1s for 10s after applying Astral Mark.

Level 3

Astral Cycle

Astral Mark

Increase the duration of Astral Mark by 1.6s.

Level 4

Luminary

Astral Mark

All allies already affected by Astral Mark are Healed for an additional 285 when you use Astral Mark.

Level 1

Frontline Commander

Increase allies and decrease enemies' Movement Speed by 35% for 4 seconds when passing through Abyssal Fortress.

Level 5

Brutal Pursuer

Rift Slash

Increase your Movement Speed by 8% for 5s after casting Rift Slash.

Level 2

Unyielding Advance

Increase Abyssal Fortress' Health by 400

Level 2

Unbreakable Will

Reduce the Health reduction from placing Abyssal Fortress >30 units away by 80%.

Level 4

Forever Changed

Increase your maximum Health by 300.

Level 2

Abyssal Breach

Enemies that pass through Abyssal Fortress are Revealed and take 10% additional weapon damage from you and your allies for 3s. Reduce Abyssal Fortress' Cooldown by 2s.

Level 1

Brave and Bold

Armor

Increase your maximum Health by 450.

Level 3

Healing Station

Turret

Heal for 100 every 1s while standing near any of your Turrets.

Level 4

Combat Repair

Turret

Heal all of your Turrets for 180 every 1s while you are standing near them.

Level 2

Bowling Ball

Rocket Boots

Gain a 300-Health Shield for 3s after activating Rocket Boots.

Level 2

Bunker

Barricade

Increase the Health of Barricade by 800.

Level 4

Fortify

Barricade

Increase the maximum Health of Barricade by 2000 and reduce its Cooldown by 3s.

Level 1

Fanning the Flames

Juggernaut

Gain a 300-Health Shield for 3s after activating Juggernaut.

Level 2

Subjugation

Juggernaut

Increase the distance traveled by Juggernaut by 8%.

Level 1

Sinister Allies

Armor

Increase your maximum Health by 750.

Level 5

Desperation

Armor

Gain a 1000-Health Shield for 3s after dropping to or below 30% Health.

Level 4

Shattered Essence

Soul Harvest

Each Soul Fragment gathered Heals you for 45.

Level 3

Earthsplitter

Cataclysm

Increase your Ultimate charge rate by 60%.

Level 1