Perennial

Blossom

Reduce the Cooldown of Blossom by 1.6s.

Level 4

Chop Down

Crippling Throw

Reduce the Cooldown of Crippling Throw by 0.6s.

Level 1

Verdant Expanse

Blossom

Increase the radius of Blossom by 16%.

Level 4

Unstoppable

Blossom

Reduce your damage taken by 24% for 4s after using Blossom.

Level 4

Fatalis

Crippling Throw

Hitting an enemy with Crippling Throw reduces the Cooldown of Blossom by 1.2s.

Level 2

Deep Roots

Crippling Throw

Crippling Throw chains to nearby enemies up to 2 times, causing a Healing aura to manifest around the nearby enemies and the original one, Healing nearby allies for 50 every 0.5s. Remedy (+10% Healing +5% Damage)

Level 1

Scribe's Wit

Nullify

Reduce the Cooldown of Nullify by 5s.

Level 5

Arcane Etching

Nullify

Gain a 100-Health Shield after activating Nullify.

Level 1

Glyph of Freedom

Armor

Reduce your active Cooldowns by 24% after getting an Elimination.

Level 3

Glyph of Health

Armor

Increase your maximum Health by 750.

Level 5

Eldritch Speed

Recharge

Increase your Movement Speed by 10% during Recharge and for 3s after it ends.

Level 1

Direct Current

Nullify

Double the range at which you can use Nullify.

Level 1

Heavy Metal

Armor

Increase your maximum Health by 750.

Level 5

Trebuchet

Siege Shield

Generate 2 Ammo after activating Siege Shield.

Level 1

Vanguard

Shoulder Bash

Reduce your damage taken by 24% for 2s after using Shoulder Bash.

Level 3

Indomitable

Armor

Gain 50% Lifesteal.

Level 5

Fervor

Armor

Increase your maximum Ammo by 1.

Level 1

Fortress Breaker

Siege Shield

Increase Siege Shield's maximum Health by 2500 and its size by 50%, but reduce its Movement Speed by 80%.

Level 1

Bragging Rights

Weapon

Hitting enemies with Bladed Chakrams Heals you for 75.

Level 5

Crowd-Pleaser

Combat Trance

Generate 3 Ammo after activating Combat Trance.

Level 3

Instrument of Fate

Heavy Blade

Reduce the Cooldown of Combat Trance by 0.8s for each enemy hit by Heavy Blade.

Level 2

World-traveler

Crouching Tigron

Reset the Cooldown of Crouching Tigron after falling to or below 30% Health.

Level 2

Flying Chakrams

Weapon

Increase your maximum Ammo by 3.

Level 3

Predatory Instincts

Weapon

Bladed Chakrams are able to bounce 2 additional times before disappearing.

Level 1

Heavy Metal

Armor

Increase your maximum Health by 750.

Level 5

Trebuchet

Siege Shield

Generate 2 Ammo after activating Siege Shield.

Level 1

Vanguard

Shoulder Bash

Reduce your damage taken by 24% for 2s after using Shoulder Bash.

Level 3

Indomitable

Armor

Gain 50% Lifesteal.

Level 5

Fervor

Armor

Increase your maximum Ammo by 1.

Level 1

Fortress Breaker

Siege Shield

Increase Siege Shield's maximum Health by 2500 and its size by 50%, but reduce its Movement Speed by 80%.

Level 1

Moonwalk

Armor

Increase your maximum Moonlight capacity by 40%.

Level 5

Protectors

Guardian Spirit

Regenerate 10% Moonlight every 1s while within 30 units of Luna and not using Moonlight.

Level 5

Broken Deity

Guardian Spirit

Increase Luna's deploy range by 40%.

Level 2

Spirit Arrows

Moonlight

Reduce the Cooldown of Luna's redeploy by 0.2s after healing with Moonlight.

Level 2

Moonlight Garden

Weapon

Generate 2% Moonlight for each enemy hit with Light Bow.

Level 1

Sacrifice

Guardian Spirit

[Guardian Spirit] If Io falls below 200 Health while Luna is active, she trades places with Luna, Heals for 50% of her Health, and Luna dies in her stead.

Level 1

Brittle

Shatter

Reduce the Cooldown of Shatter by 2.5s.

Level 5

Squadron

Illusion

Increase the Health of each of your Illusions by 300.

Level 3

Spring Bloom

Illusion

Reduce the Cooldown of Illusion by 1.5s when an Illusion dies to an enemy.

Level 3

Fracture

Shatter

Heal your Illusions for 40% of their maximum Health after activating Shatter.

Level 2

Carry On

Illusion

Increase the duration of your Illusions by 2s.

Level 2

Life Exchange

Shatter

Shatter instantly Heals your target for 800, but its Cooldown is increased by 1s and it no longer explodes your Illusions. Reduce the Cooldown of Shatter by 2s if you miss.

Level 1

Moonwalk

Armor

Increase your maximum Moonlight capacity by 40%.

Level 5

Protectors

Guardian Spirit

Regenerate 10% Moonlight every 1s while within 30 units of Luna and not using Moonlight.

Level 5

Broken Deity

Guardian Spirit

Increase Luna's deploy range by 40%.

Level 2

Spirit Arrows

Moonlight

Reduce the Cooldown of Luna's redeploy by 0.2s after healing with Moonlight.

Level 2

Moonlight Garden

Weapon

Generate 2% Moonlight for each enemy hit with Light Bow.

Level 1

Sacrifice

Guardian Spirit

[Guardian Spirit] If Io falls below 200 Health while Luna is active, she trades places with Luna, Heals for 50% of her Health, and Luna dies in her stead.

Level 1

Raze and Burn

Piercing Quills

Heal for 200 after hitting an enemy with Piercing Quills.

Level 4

Collateral Damage

Form Swap

Gain 10% Lifesteal with Caustic Spray for 3s after swapping into Planted Form.

Level 1

Futile Efforts

Weapon

Reduce the Cooldown of Piercing Quills by 0.15000000000000002s each time you hit an enemy with Caustic Spray.

Level 3

Deadly Predator

Weapon

Increase your maximum acid capacity by 32%.

Level 4

Ancient Wounds

Piercing Quills

Generate 6% of your maximum acid after hitting an enemy with Piercing Quills.

Level 3

Corrosive Acid

Caustic Spray

Augment your acid, increasing the damage done by Caustic Spray to 42, the damage of acid pools to 40, and increase the Movement Speed of allies within your acid pools by 20%.

Level 1

Dampener

Armor

Increase your maximum Health by 750.

Level 5

Refraction

Emitter

Reduce the Cooldown of Emitter by 1s.

Level 1

Crystal Capacitor

Missile Launcher

Reduce the Cooldown of Missile Launcher by 0.7s.

Level 1

Fuel Reserves

Advance

Increase the travel distance of Advance by 28%.

Level 4

Metal March

Missile Launcher

Reduce your damage taken by 12% for 2s after using Missile Launcher.

Level 4

Rocket Barrage

Missile Launcher

Increase the radius of Missile Launcher's explosions by 66%.

Level 1

Harbinger

Soul Harvest

Reduce the Cooldown of Juggernaut by 1s after activating Soul Harvest.

Level 1

Sinister Allies

Armor

Increase your maximum Health by 750.

Level 5

Shattered Essence

Soul Harvest

Each Soul Fragment gathered Heals you for 60.

Level 4

Void Lord

Soul Harvest

Reduce your damage taken by 20% for 3s after activating Soul Harvest.

Level 4

Tormented Fissure

Ignition

Generate 25 Ammo after activating Ignition.

Level 1

Subservience

Soul Harvest

All your living allies are Healed for 1.5% of their maximum Health for each Soul Harvested.

Level 1

Mortal Skewer

Harpoon

Heal for 350 for each enemy hit with Harpoon.

Level 5

Dreadnought

Armor

Increase your maximum Health by 50.

Level 1

Expansive Vault

Shortcut

Jump height is increased for your first jump within 1s after exiting a Shortcut.

Level 1

Dark Bargain

Shortcut

Heal for 250 every 1s while standing near a Shortcut.

Level 5

Cursed Weaponry

Harpoon

Generate 3 Ammo for each enemy hit by Harpoon.

Level 3

Hurl

Harpoon

Harpoon no longer has a Cooldown, but no longer Slows enemies hit.

Level 1

Surprise!

Shadow Step

Generate 20 Ammo after activating Shadow Step.

Level 4

Trigger Happy

Weapon

Increase your Healing received by 15% while firing your SMGs.

Level 3

Tenacious

Shadow Step

Heal for 200 after activating Shadow Step.

Level 5

Memento

Weapon

Increase your maximum Ammo by 10.

Level 2

Harsh Training

Armor

Increase your maximum Health by 50.

Level 1

Level 1

Moonwalk

Armor

Increase your maximum Moonlight capacity by 40%.

Level 5

Protectors

Guardian Spirit

Regenerate 10% Moonlight every 1s while within 30 units of Luna and not using Moonlight.

Level 5

Broken Deity

Guardian Spirit

Increase Luna's deploy range by 40%.

Level 2

Spirit Arrows

Moonlight

Reduce the Cooldown of Luna's redeploy by 0.2s after healing with Moonlight.

Level 2

Moonlight Garden

Weapon

Generate 2% Moonlight for each enemy hit with Light Bow.

Level 1

Sacrifice

Guardian Spirit

[Guardian Spirit] If Io falls below 200 Health while Luna is active, she trades places with Luna, Heals for 50% of her Health, and Luna dies in her stead.

Level 1