Reduce the Cooldown of Hardening by 0.6s after hitting an enemy with Piercing Quills.
Level 2
Ancient Wounds
Piercing Quills
Generate 6% of your maximum acid after hitting an enemy with Piercing Quills.
Level 3
End of an Era
Hardening
Increase the additional Damage Reduction benefit of Hardening by 80% but reduce its duration by {scale=10|10}%.
Level 4
Extermination
Armor
Reduce the duration and effectiveness of Crowd Control by 25%.
Level 5
Sight Begets Strength
Primal Vision
Allied players within range of your Primal Vision gain a stackable 90-Health Shield and 15% increased Movement Speed for 4s each time you pulse, but your Primal Vision has its Cooldown increased by 2s.