Apex Predator

Armor

Increase your maximum Health by 200.

Level 4

Altitude

Armor

Restore 20% of your maximum Booster fuel after getting a Killing Blow or Elimination.

Level 1

Masterful

Fire Spit

Reduce the Cooldown of Fire Spit by 2s for each enemy hit by it.

Level 4

Rain of Fire

Thrust

Consume 60% less Booster fuel for 5s after using Thrust.

Level 4

Condescension

Fire Spit

Reduce the Cooldown of Thrust by 2.4s for each enemy hit with Fire Spit.

Level 2

Fusillade

Weapon

Damage dealt when hitting an enemy directly with Rocket Launcher or Salvo is increased by 25%.

Level 1

Depths of Despair

Reduce the damage you take by 12% for 2s after activating Conviction.

Level 3

Tools of Salvation

Increase your maximum Health by 300.

Level 2

Ad Eternum

Reduce the damage you take by 6% while at least one wall segment of Sanctuary is active.

Level 2

Grim Deliverance

Heal for 25 for every percentage point of Ire consumed by abilities.

Level 5

Forged in Battle

Reduce the Cooldown of Sanctuary by 1.7999999999999998s.

Level 3

Persistence

Gain 30% Lifesteal on your hits with Judgement.

Level 1

Depths of Despair

Reduce the damage you take by 12% for 2s after activating Conviction.

Level 3

Tools of Salvation

Increase your maximum Health by 300.

Level 2

Ad Eternum

Reduce the damage you take by 6% while at least one wall segment of Sanctuary is active.

Level 2

Grim Deliverance

Heal for 25 for every percentage point of Ire consumed by abilities.

Level 5

Forged in Battle

Reduce the Cooldown of Sanctuary by 1.7999999999999998s.

Level 3

Persistence

Gain 30% Lifesteal on your hits with Judgement.

Level 1

Dance with Fate

Tendril

Reduce the cooldown of Tendril by 2.4s.

Level 4

Unified in Purpose

Dark Siphon

Reduce the damage you take by 10% for 3s after hitting an enemy with Dark Siphon.

Level 3

Elegant Paring

Tendril

Reduce the Cooldown of Dark Siphon and Obliteration by 0.8s after activating Tendril.

Level 2

What May Come

Tendril

Reduce the damage you take by 16% for 3s after activating Tendril.

Level 4

A Change of Faith

Dark Siphon

Reduce the Cooldown of Tendril by 1.2s after hitting an enemy with Dark Siphon.

Level 2

Relentless Presence

Tendril

Tendril gains a second charge, but its Cooldown is increased by 2s.

Level 1

Dance with Fate

Tendril

Reduce the cooldown of Tendril by 2.4s.

Level 4

Unified in Purpose

Dark Siphon

Reduce the damage you take by 10% for 3s after hitting an enemy with Dark Siphon.

Level 3

Elegant Paring

Tendril

Reduce the Cooldown of Dark Siphon and Obliteration by 0.8s after activating Tendril.

Level 2

What May Come

Tendril

Reduce the damage you take by 16% for 3s after activating Tendril.

Level 4

A Change of Faith

Dark Siphon

Reduce the Cooldown of Tendril by 1.2s after hitting an enemy with Dark Siphon.

Level 2

Relentless Presence

Tendril

Tendril gains a second charge, but its Cooldown is increased by 2s.

Level 1

Broken Deity

Guardian Spirit

Increase Luna's deploy range by 60%.

Level 3

Sky-Walker

Armor

Lower your gravity and increase your air control by 20% while firing your Light Bow.

Level 2

Celestial Body

Armor

Increase your maximum Health by 100.

Level 2

Full Moon

Lunar Leap

Gain a 300-Health Shield for 3s after activating Lunar Leap.

Level 3

Lunar Connection

Guardian Spirit

Reduce the cooldown of Luna's initial deploy by 6s.

Level 5

Sacrifice

Guardian Spirit

[Guardian Spirit] If Io falls below 200 Health while Luna is active, she trades places with Luna, Heals for 50% of her Health, and Luna dies in her stead.

Level 1

Broken Deity

Guardian Spirit

Increase Luna's deploy range by 60%.

Level 3

Sky-Walker

Armor

Lower your gravity and increase your air control by 20% while firing your Light Bow.

Level 2

Celestial Body

Armor

Increase your maximum Health by 100.

Level 2

Full Moon

Lunar Leap

Gain a 300-Health Shield for 3s after activating Lunar Leap.

Level 3

Lunar Connection

Guardian Spirit

Reduce the cooldown of Luna's initial deploy by 6s.

Level 5

Sacrifice

Guardian Spirit

[Guardian Spirit] If Io falls below 200 Health while Luna is active, she trades places with Luna, Heals for 50% of her Health, and Luna dies in her stead.

Level 1

Dance with Fate

Tendril

Reduce the cooldown of Tendril by 2.4s.

Level 4

Unified in Purpose

Dark Siphon

Reduce the damage you take by 10% for 3s after hitting an enemy with Dark Siphon.

Level 3

Elegant Paring

Tendril

Reduce the Cooldown of Dark Siphon and Obliteration by 0.8s after activating Tendril.

Level 2

What May Come

Tendril

Reduce the damage you take by 16% for 3s after activating Tendril.

Level 4

A Change of Faith

Dark Siphon

Reduce the Cooldown of Tendril by 1.2s after hitting an enemy with Dark Siphon.

Level 2

Relentless Presence

Tendril

Tendril gains a second charge, but its Cooldown is increased by 2s.

Level 1

Level 5

Level 4

Level 1

Level 3

Level 2

Level 1

Wings of Terror

Death Wings

Increase the lateral movement of Death Wings by 12%.

Level 2

Unholy Flight

Death Wings

Reduce your damage taken by 20% for 1.5s after activating Death Wings.

Level 4

Blood Cannon

Armor

Increase extra Blood Health by 400 while decreasing base Health by {scale=125|125}.

Level 2

Tainted Form

Weapon

Gain 5% of Max Health when hitting a marked enemy.

Level 5

Gloomy Nook

Armor

Gain a 300 Shield for 1.75s after falling to or below 75% Health. Can only happen once every 35s.

Level 2

Cursed Accord

Armor

Decrease the Blood cost of your abilities by 35% and increase your Healing done by 15%, but reduce the damage you do by 25%.

Level 1

Bushwhack

Quick Switch

Swapping to your Pistol grants 50% Lifesteal for 3s.

Level 5

Roost

Scope

Hits with Talon Rifle generate 3% Ultimate charge.

Level 3

Escape Plan

Stealth

Heal for 500 after entering Stealth at or below 65% Health.

Level 5

Infused Crystals

Weapon

Increase the maximum Ammo count of both your weapons by 1.

Level 1

Cooled Mags

Quick Switch

Your inactive weapon generates 2 Ammo every 2s.

Level 1

Unauthorized Use

Flare

Flare deals 250% base damage, has significantly increased Projectile Speed, and has its Cooldown reduced by 10s, but no longer Reveals enemies.

Level 1

Broken Deity

Guardian Spirit

Increase Luna's deploy range by 60%.

Level 3

Sky-Walker

Armor

Lower your gravity and increase your air control by 20% while firing your Light Bow.

Level 2

Celestial Body

Armor

Increase your maximum Health by 100.

Level 2

Full Moon

Lunar Leap

Gain a 300-Health Shield for 3s after activating Lunar Leap.

Level 3

Lunar Connection

Guardian Spirit

Reduce the cooldown of Luna's initial deploy by 6s.

Level 5

Sacrifice

Guardian Spirit

[Guardian Spirit] If Io falls below 200 Health while Luna is active, she trades places with Luna, Heals for 50% of her Health, and Luna dies in her stead.

Level 1

Broken Deity

Guardian Spirit

Increase Luna's deploy range by 60%.

Level 3

Sky-Walker

Armor

Lower your gravity and increase your air control by 20% while firing your Light Bow.

Level 2

Celestial Body

Armor

Increase your maximum Health by 100.

Level 2

Full Moon

Lunar Leap

Gain a 300-Health Shield for 3s after activating Lunar Leap.

Level 3

Lunar Connection

Guardian Spirit

Reduce the cooldown of Luna's initial deploy by 6s.

Level 5

Sacrifice

Guardian Spirit

[Guardian Spirit] If Io falls below 200 Health while Luna is active, she trades places with Luna, Heals for 50% of her Health, and Luna dies in her stead.

Level 1