Run and Gun

Hustle

Regenerate 4 Ammo every 1s while using Hustle.

Level 2

Scramble

Hustle

Reduce damage taken by 12% while using Hustle.

Level 4

Hot Potato

Frag Grenade

Reduce the cook time of Frag Grenade by 20%.

Level 2

Second Wind

Hustle

Increase your Movement Speed after deactivating Hustle by 30% for 1.5s.

Level 3

Guerrilla Warfare

Hustle

Reduce the Cooldown of Frag Grenade by 0.8s every 1s while Hustle is active.

Level 4

Burst Mode

Weapon

Switch your rifle to Three-Shot-Burst mode, increasing your effective range and firing three 210-damage shots every 0.4s.

Level 1

The Great Worm

Form Swap

Gain a 240-Health Shield for 2s after swapping into Travel Form.

Level 3

Violent Birth

Hardening

Reduce the Cooldown of Acceleration by 1.6s after activating Hardening.

Level 2

Triumphant Return

Form Swap

Reduce your acid consumption by 90% for 0.8s after swapping into Planted Form.

Level 2

Deadly Pursuit

Form Swap

Increase your Movement Speed by 20% for 3s after swapping into Travel Form.

Level 3

Towering Presence

Armor

Increase the Health of your Ultimate Health pool by 1000.

Level 5

Corrosive Acid

Caustic Spray

Augment your acid, increasing the damage done by Caustic Spray to 42, the damage of acid pools to 40, and increase the Movement Speed of allies within your acid pools by 20%.

Level 1

Deadly Prey

Increase your Movement Speed by 40% for 2s after Body and Soul ends.

Level 4

Sealed Fate

Heal for 140 after an enemy survives being hit with Yokai Doll or Savage Tear at maximum Curse stacks.

Level 4

Sudden Death

Increase the duration of Curse stacks by 1.2s.

Level 2

Cursed Doll

Apply 3 stacks of Curse to the target hit by your Savage Tear. This effect can occur once every -20s.

Level 4

Folk Tales

Increase your maximum Health by 50.

Level 1

Empowered Curse

Increase your Ultimate charge rate by 3% and Movement Speed by 5% per stack of Curse currently on enemies.

Level 1

Trigger Happy

Weapon

Increase your Healing received by 15% while firing your SMGs.

Level 3

Surprise!

Shadow Step

Generate 15 Ammo after activating Shadow Step.

Level 3

Tenacious

Shadow Step

Heal for 160 after activating Shadow Step.

Level 4

Trespasser

Agility

Increase your jump height by 20% while Agility is active.

Level 1

Memento

Weapon

Increase your maximum Ammo by 20.

Level 4

Adrenaline Junkie

Weapon

Generate 1 Energy for every 80 damage done with your Submachine Guns.

Level 1

Astral Traveler

Ghost Walk

Increase your Movement Speed while using Ghost Walk by 20%.

Level 2

Outreach

Healing Totem

Increase the deploy range of Healing Totem by 75%.

Level 5

Haunting

Ghost Walk

Reduce the Cooldown of Ghost Walk by 1.2s.

Level 2

Gale

Ghost Walk

Decrease the Cooldown of Healing Totem by 3s after activating Ghost Walk.

Level 5

Crackle

Healing Totem

Increase the duration of Healing Totem by 0.2s.

Level 1

Totemic Ward

Healing Totem

Gain a third charge of Healing Totem, increase their healing by 15%, and increase their radius of 20%. Remedy(+10% Healing and +5% Damage)

Level 1

Hellish Lodestones

Ignition

Reduce your damage taken by 12% for 3s after activating Ignition.

Level 4

Sinister Allies

Armor

Increase your maximum Health by 150.

Level 1

Harbinger

Soul Harvest

Reduce the Cooldown of Juggernaut by 4s after activating Soul Harvest.

Level 4

Void Lord

Soul Harvest

Reduce your damage taken by 20% for 3s after activating Soul Harvest.

Level 4

Subjugation

Juggernaut

Increase the distance traveled by Juggernaut by 16%.

Level 2

Enforcer

Juggernaut

Reduce your damage taken by 40% while using Juggernaut.

Level 1

Hulking Monstrosity

Armor

Increase your maximum Health by 450.

Level 3

Powerslave

Power Siphon

Reduce the Movement Speed penalty during Power Siphon by 15%.

Level 1

Strength of Stone

Calamity Blast

Reduce your damage taken by up to 16% based on the number of Calamity Blast charges you have stored.

Level 4

It Follows

Calamity Blast

Increase your Movement Speed by 10% for each Calamity Charge you have.

Level 5

It Watches

Armor

Generate 6% Power Siphon charge for each enemy hit with Massacre Axe.

Level 2

Undying

Armor

Reduce the damage you take by 20% while at or below 50% Health.

Level 1

Latch and Fire

Explosive Dodge

Reduce the Cooldown of Grappling Hook by 1s after activating Explosive Dodge.

Level 2

The Fear

Trigger Discipline

Reduce your active Cooldowns by 4s after getting a Killing Blow while in Burst Mode.

Level 4

Night Watch

Trigger Discipline

Generate 8 Ammo when you activate Trigger Discipline.

Level 2

Never Look

Armor

Increase your maximum Health by 200.

Level 4

Red Room

Trigger Discipline

Increase your Movement Speed by 12% while in Automatic fire mode.

Level 3

Tribunal Upgrades

Heavy SMG has three elemental effects: Fire imbued Mag Dump -- dealing damage over time, Electrically charged burst mode -- dealing AOE damage on stacks, and Ice infused Automatic -- shots slows enemies.

Level 1

Watchful

Armor

Reduce all active Cooldowns by 20% after getting an Elimination.

Level 2

Vengeance

Defiance

Heal for 120 after hitting at least one enemy with Defiance.

Level 4

Marksman

Defiance

Generate 2 Ammo after hitting an enemy with Defiance.

Level 2

Abyssal Touch

Defiance

Reduce the Cooldown of Nether Step by 6s after hitting an enemy with Defiance.

Level 5

Elusive

Nether Step

Increase your Movement Speed by 20% for 3s after activating Nether Step.

Level 2

Defiant Fist

Defiance

Successful hits with your Revolver increase the damage of your next Defiance by 20%, stacking up to 100%.

Level 1

Crow's Nest

Broadside

Hitting yourself with Broadside now applies a Knockback of 300.

Level 2

Expansive Vault

Shortcut

Jump height is increased for your first jump within 1s after exiting a Shortcut.

Level 1

Hangman's Ire

Armor

Gain a 600-Health Shield after dropping to or below 50% Health.

Level 4

Dark Bargain

Shortcut

Heal for 200 every 1s while standing near a Shortcut.

Level 4

Sea Dogs

Armor

Increase the Healing you receive by 20% while at or below 50% Health.

Level 4

Scuttle

Weapon

The projectile launched by reloading your Cursed Howitzer explodes in a 20-unit-radius area on contact.

Level 1

Salvage

Dredge Anchor

Regenerate 1 Ammo every 0.5s for 0.5s after activating Dredge Anchor.

Level 1

Surf

Shell Spin

Increase the travel distance of Shell Spin by 30%.

Level 3

Ebb and Flow

Armor

Reduce the Cooldown of Shell Spin by 60% after falling to or below 40% Health.

Level 3

Harden

Dredge Anchor

Reduce your damage taken by 15% for 2s after hitting an enemy with Dredge Anchor.

Level 3

Strongarm

Dredge Anchor

Reduce the Cooldown of Dredge Anchor by 3s.

Level 5

Pluck

Dredge Anchor

For 4s after hitting an enemy with Dredge Anchor, your next weapon shot against that enemy deals 75% increased damage.

Level 1

Lifetaker

Armor

Reduce your active Cooldowns by 8% after getting an Elimination.

Level 1

Into the Breach!

Battle Shout

Increase your Movement Speed by 32% for 3s after activating Battle Shout.

Level 4

Excessive Force

Weapon

Increase your maximum Ammo by 3.

Level 1

Platemail

Armor

Increase your maximum Health by 600.

Level 4

Never Surrender!

Battle Shout

Reduce the Cooldown of Battle Shout by 3s.

Level 5

Lian's Shield

Bulwark

Your Shield now regenerates at 120% effectiveness, and regenerates even while it is active.

Level 1

Opulence

Missile Launcher

Heal 240 health for each player hit by Missile Launcher.

Level 4

No Chill

Weapon

Increase your Movement Speed while firing by 10%.

Level 2

Proximity

Advance

Reduce your damage taken by 3% for 3s after activating Advance.

Level 1

Crystal Capacitor

Missile Launcher

Reduce the Cooldown of Missile Launcher by 2.8s.

Level 4

Metal March

Missile Launcher

Reduce your damage taken by 12% for 2s after using Missile Launcher.

Level 4

Rocket Barrage

Missile Launcher

Increase the radius of Missile Launcher's explosions by 66%.

Level 1

Fatalis

Crippling Throw

Hitting an enemy with Crippling Throw reduces the Cooldown of Blossom by 1.7999999999999998s.

Level 3

Force of Nature

Armor

Gain a 300-Health Shield for 3s after dropping to or below 50% Health.

Level 2

Chopper

Crippling Throw

Heal for 150 after hitting an enemy with Crippling Throw.

Level 3

Barkskin

Armor

Increase your maximum Health by 150.

Level 2

Chop Down

Crippling Throw

Reduce the Cooldown of Crippling Throw by 3s.

Level 5

Ferocity

Weapon

Increase your maximum damage scaling over distance by 60%. Surgery (+10% Damage)

Level 1

Combat Repair

Turret

Heal all of your Turrets for 180 every 1s while you are standing near them.

Level 2

Brave and Bold

Armor

Increase your maximum Health by 300.

Level 2

One Man's Scrap

Armor

When a Turret is destroyed, reduce your active Cooldowns by 25%.

Level 5

Field Deploy

Turret

Heal for 60 each time one of your base Turrets hits an enemy.

Level 3

Healing Station

Turret

Heal for 75 every 1s while standing near any of your Turrets.

Level 3

Forgefire

Dome Shield

Dome Shield now costs 40% Ultimate charge and places a Flamethrower Turret with modified values and no Shield. Can have two active at once.

Level 1

Level 1

Level 4

Level 3

Level 2

Level 5

Level 1

Follow the Scent

Fire Spit

Increase the size of your Fire Spit by 48%.

Level 2

Rain of Fire

Thrust

Consume 30% less Booster fuel for 5s after using Thrust.

Level 2

Condescension

Fire Spit

Reduce the Cooldown of Thrust by 2.4s for each enemy hit with Fire Spit.

Level 2

Masterful

Fire Spit

Reduce the Cooldown of Fire Spit by 2.5s for each enemy hit by it.

Level 5

Lung Capacity

Fire Spit

Heal for 200 for each enemy hit with Fire Spit.

Level 4

Combustible

Fire Spit

Fire Spit applies a Knockback and deals an additional 300 damage over 2s.

Level 1

Backdraft

Detonate

Reduce the Cooldown of Poppy Bomb by 0.1s for each Sticky Bomb you Detonate.

Level 1

Royal Decree

Detonate

Increase your Reload Speed by 16% for 3s after using Detonate.

Level 4

King's New Cloak

Armor

Increase your maximum Health by 250.

Level 5

King's Court

Detonate

Heal for 60 for each Sticky Bomb you Detonate.

Level 4

Countdown

Grumpy Bomb

Reduce the Cooldown of Grumpy Bomb by 0.8s.

Level 1

Chain Reaction

Detonate

Hitting with more than one Sticky Bomb on a use of Detonate increases the damage of each explosion after the first by 30% of its base damage, up to 150%.

Level 1

Level 3

Level 5

Level 2

Level 2

Level 3

Level 1