Heal for 30 every time you Heal an ally with Moonlight.
Level 3
Celestial Body
Armor
Increase your maximum Health by 200.
Level 4
Feral Strength
Guardian Spirit
Increase Luna's Health by 225.
Level 1
Eternal Reliquary
Lunar Leap
Generate 4 Ammo after using Lunar Leap.
Level 2
Sacrifice
Guardian Spirit
[Guardian Spirit] If Io falls below 200 Health while Luna is active, she trades places with Luna, Heals for 50% of her Health, and Luna dies in her stead.