Reduce the Cooldown of Hardening by 0.3s after hitting an enemy with Piercing Quills.
Level 1
Raze and Burn
Piercing Quills
Heal for 100 after hitting an enemy with Piercing Quills.
Level 2
Collateral Damage
Form Swap
Gain 20% Lifesteal with Caustic Spray for 3s after swapping into Planted Form.
Level 2
Spreading Domain
Hardening
Heal for 500 if you do not cancel Hardening before it ends.
Level 5
End of an Era
Hardening
Increase the additional Damage Reduction benefit of Hardening by 100% but reduce its duration by {scale=10|10}%.
Level 5
Unnatural Persistence
Armor
Heal for 25 every 0.25s while on an objective and for 1s after leaving the objective. This effect increases the longer you maintain it, up to 65 every 0.25s.