Crescent

Lunar Leap

Reset the Cooldown of Lunar Leap after dropping to or below 30% Health.

Level 2

Half Moon

Lunar Leap

Reduce the Cooldown of Lunar Leap by 1.6s.

Level 2

Feral Strength

Guardian Spirit

Increase Luna's Health by 675.

Level 3

Full Moon

Lunar Leap

Gain a 300-Health Shield for 3s after activating Lunar Leap.

Level 3

Restored Faith

Moonlight

Heal for 50 every time you Heal an ally with Moonlight.

Level 5

Sacrifice

Guardian Spirit

[Guardian Spirit] If Io falls below 200 Health while Luna is active, she trades places with Luna, Heals for 50% of her Health, and Luna dies in her stead.

Level 1

Crescent

Lunar Leap

Reset the Cooldown of Lunar Leap after dropping to or below 30% Health.

Level 2

Half Moon

Lunar Leap

Reduce the Cooldown of Lunar Leap by 1.6s.

Level 2

Feral Strength

Guardian Spirit

Increase Luna's Health by 675.

Level 3

Full Moon

Lunar Leap

Gain a 300-Health Shield for 3s after activating Lunar Leap.

Level 3

Restored Faith

Moonlight

Heal for 50 every time you Heal an ally with Moonlight.

Level 5

Sacrifice

Guardian Spirit

[Guardian Spirit] If Io falls below 200 Health while Luna is active, she trades places with Luna, Heals for 50% of her Health, and Luna dies in her stead.

Level 1

Crescent

Lunar Leap

Reset the Cooldown of Lunar Leap after dropping to or below 30% Health.

Level 2

Half Moon

Lunar Leap

Reduce the Cooldown of Lunar Leap by 1.6s.

Level 2

Feral Strength

Guardian Spirit

Increase Luna's Health by 675.

Level 3

Full Moon

Lunar Leap

Gain a 300-Health Shield for 3s after activating Lunar Leap.

Level 3

Restored Faith

Moonlight

Heal for 50 every time you Heal an ally with Moonlight.

Level 5

Sacrifice

Guardian Spirit

[Guardian Spirit] If Io falls below 200 Health while Luna is active, she trades places with Luna, Heals for 50% of her Health, and Luna dies in her stead.

Level 1

Bragging Rights

Weapon

Hitting enemies with Bladed Chakrams Heals you for 75.

Level 5

Crowd-Pleaser

Combat Trance

Generate 3 Ammo after activating Combat Trance.

Level 3

Instrument of Fate

Heavy Blade

Reduce the Cooldown of Combat Trance by 0.8s for each enemy hit by Heavy Blade.

Level 2

World-traveler

Crouching Tigron

Reset the Cooldown of Crouching Tigron after falling to or below 30% Health.

Level 2

Flying Chakrams

Weapon

Increase your maximum Ammo by 3.

Level 3

Predatory Instincts

Weapon

Bladed Chakrams are able to bounce 2 additional times before disappearing.

Level 1

Crescent

Lunar Leap

Reset the Cooldown of Lunar Leap after dropping to or below 30% Health.

Level 2

Half Moon

Lunar Leap

Reduce the Cooldown of Lunar Leap by 1.6s.

Level 2

Feral Strength

Guardian Spirit

Increase Luna's Health by 675.

Level 3

Full Moon

Lunar Leap

Gain a 300-Health Shield for 3s after activating Lunar Leap.

Level 3

Restored Faith

Moonlight

Heal for 50 every time you Heal an ally with Moonlight.

Level 5

Sacrifice

Guardian Spirit

[Guardian Spirit] If Io falls below 200 Health while Luna is active, she trades places with Luna, Heals for 50% of her Health, and Luna dies in her stead.

Level 1

Crescent

Lunar Leap

Reset the Cooldown of Lunar Leap after dropping to or below 30% Health.

Level 2

Half Moon

Lunar Leap

Reduce the Cooldown of Lunar Leap by 1.6s.

Level 2

Feral Strength

Guardian Spirit

Increase Luna's Health by 675.

Level 3

Full Moon

Lunar Leap

Gain a 300-Health Shield for 3s after activating Lunar Leap.

Level 3

Restored Faith

Moonlight

Heal for 50 every time you Heal an ally with Moonlight.

Level 5

Sacrifice

Guardian Spirit

[Guardian Spirit] If Io falls below 200 Health while Luna is active, she trades places with Luna, Heals for 50% of her Health, and Luna dies in her stead.

Level 1

Inevitability

Obliteration

Reduce the Cooldown of Dark Siphon by 0.8s after hitting at least one enemy with Obliteration.

Level 2

A Broken Path

Dark Siphon

Increase the Healing received from Dark Siphon by 200 health.

Level 4

New Purpose

Armor

Increase your maximum Health by 50.

Level 1

Unified in Purpose

Dark Siphon

Reduce the damage you take by 10% for 3s after hitting an enemy with Dark Siphon.

Level 3

Crimson Ascent

Dark Siphon

Reduce the Cooldown of Dark Siphon by 5s if it fails to hit a target.

Level 5

Unyielding Pressure

Your Dark Siphon now deals an additional 10% of the target's maximum Health as damage over 1.5s.

Level 1

Crescent

Lunar Leap

Reset the Cooldown of Lunar Leap after dropping to or below 30% Health.

Level 2

Half Moon

Lunar Leap

Reduce the Cooldown of Lunar Leap by 1.6s.

Level 2

Feral Strength

Guardian Spirit

Increase Luna's Health by 675.

Level 3

Full Moon

Lunar Leap

Gain a 300-Health Shield for 3s after activating Lunar Leap.

Level 3

Restored Faith

Moonlight

Heal for 50 every time you Heal an ally with Moonlight.

Level 5

Sacrifice

Guardian Spirit

[Guardian Spirit] If Io falls below 200 Health while Luna is active, she trades places with Luna, Heals for 50% of her Health, and Luna dies in her stead.

Level 1

Crescent

Lunar Leap

Reset the Cooldown of Lunar Leap after dropping to or below 30% Health.

Level 2

Half Moon

Lunar Leap

Reduce the Cooldown of Lunar Leap by 1.6s.

Level 2

Feral Strength

Guardian Spirit

Increase Luna's Health by 675.

Level 3

Full Moon

Lunar Leap

Gain a 300-Health Shield for 3s after activating Lunar Leap.

Level 3

Restored Faith

Moonlight

Heal for 50 every time you Heal an ally with Moonlight.

Level 5

Sacrifice

Guardian Spirit

[Guardian Spirit] If Io falls below 200 Health while Luna is active, she trades places with Luna, Heals for 50% of her Health, and Luna dies in her stead.

Level 1

Broken Deity

Guardian Spirit

Increase Luna's deploy range by 20%.

Level 1

Sky-Walker

Armor

Lower your gravity and increase your air control by 30% while firing your Light Bow.

Level 3

Half Moon

Lunar Leap

Reduce the Cooldown of Lunar Leap by 4s.

Level 5

Full Moon

Lunar Leap

Gain a 300-Health Shield for 3s after activating Lunar Leap.

Level 3

Crescent

Lunar Leap

Reset the Cooldown of Lunar Leap after dropping to or below 45% Health.

Level 3

Sacrifice

Guardian Spirit

[Guardian Spirit] If Io falls below 200 Health while Luna is active, she trades places with Luna, Heals for 50% of her Health, and Luna dies in her stead.

Level 1