Feral Strength

Guardian Spirit

Increase Luna's Health by 900.

Level 4

Swift Arrows

Moonlight

Increase allies' Movement Speed by 18% while they are being Healed by Moonlight.

Level 3

Moonwalk

Armor

Increase your maximum Moonlight capacity by 24%.

Level 3

Protectors

Guardian Spirit

Regenerate 4% Moonlight every 1s while within 30 units of Luna and not using Moonlight.

Level 2

Broken Deity

Guardian Spirit

Increase Luna's deploy range by 60%.

Level 3

Life Link

Guardian Spirit

[Guardian Spirit] Luna Heals allies around her for 320 per second. Reduce Moonlight's Healing to 135 every 0.15s.

Level 1

Mystic Fire

Frostfire Glide

Regenerate 15% Mana every 1s while using FrostFire Glide.

Level 3

Ice in her Veins

Frostfire Glide

Heal for 200 every 1s while using FrostFire Glide.

Level 4

Pyretic Momentum

Frostfire Glide

FrostFire Glide travels 10% faster.

Level 2

Pilgrimage

Frostfire Glide

Increase the duration of FrostFire Glide by 1.5s.

Level 3

Ancient Duty

Armor

Increase your maximum Health by 150.

Level 3

Pyromania

Pyre Ball

Each of your Pyre Balls explodes in a 20-unit-radius area on hit and generate more Mana, but the damage of Pyre Ball when fully charged is reduced to 800.

Level 1

Mystic Fire

Frostfire Glide

Regenerate 15% Mana every 1s while using FrostFire Glide.

Level 3

Ice in her Veins

Frostfire Glide

Heal for 200 every 1s while using FrostFire Glide.

Level 4

Pyretic Momentum

Frostfire Glide

FrostFire Glide travels 10% faster.

Level 2

Pilgrimage

Frostfire Glide

Increase the duration of FrostFire Glide by 1.5s.

Level 3

Ancient Duty

Armor

Increase your maximum Health by 150.

Level 3

Pyromania

Pyre Ball

Each of your Pyre Balls explodes in a 20-unit-radius area on hit and generate more Mana, but the damage of Pyre Ball when fully charged is reduced to 800.

Level 1

Feral Strength

Guardian Spirit

Increase Luna's Health by 900.

Level 4

Swift Arrows

Moonlight

Increase allies' Movement Speed by 18% while they are being Healed by Moonlight.

Level 3

Moonwalk

Armor

Increase your maximum Moonlight capacity by 24%.

Level 3

Protectors

Guardian Spirit

Regenerate 4% Moonlight every 1s while within 30 units of Luna and not using Moonlight.

Level 2

Broken Deity

Guardian Spirit

Increase Luna's deploy range by 60%.

Level 3

Life Link

Guardian Spirit

[Guardian Spirit] Luna Heals allies around her for 320 per second. Reduce Moonlight's Healing to 135 every 0.15s.

Level 1

Feral Strength

Guardian Spirit

Increase Luna's Health by 900.

Level 4

Swift Arrows

Moonlight

Increase allies' Movement Speed by 18% while they are being Healed by Moonlight.

Level 3

Moonwalk

Armor

Increase your maximum Moonlight capacity by 24%.

Level 3

Protectors

Guardian Spirit

Regenerate 4% Moonlight every 1s while within 30 units of Luna and not using Moonlight.

Level 2

Broken Deity

Guardian Spirit

Increase Luna's deploy range by 60%.

Level 3

Life Link

Guardian Spirit

[Guardian Spirit] Luna Heals allies around her for 320 per second. Reduce Moonlight's Healing to 135 every 0.15s.

Level 1

Out of Time

Hardening

Reduce the additional Damage Reduction given by Hardening by half. For 1.5s after using Hardening, gain its benefits.

Level 3

Collateral Damage

Form Swap

Gain 30% Lifesteal with Caustic Spray for 3s after swapping into Planted Form.

Level 3

The Great Worm

Form Swap

Gain a 160-Health Shield for 2s after swapping into Travel Form.

Level 2

Raze and Burn

Piercing Quills

Heal for 100 after hitting an enemy with Piercing Quills.

Level 2

Deadly Predator

Weapon

Increase your maximum acid capacity by 40%.

Level 5

Sight Begets Strength

Primal Vision

Allied players within range of your Primal Vision gain a stackable 90-Health Shield and 15% increased Movement Speed for 4s each time you pulse, but your Primal Vision has its Cooldown increased by 2s.

Level 1

Feral Strength

Guardian Spirit

Increase Luna's Health by 900.

Level 4

Swift Arrows

Moonlight

Increase allies' Movement Speed by 18% while they are being Healed by Moonlight.

Level 3

Moonwalk

Armor

Increase your maximum Moonlight capacity by 24%.

Level 3

Protectors

Guardian Spirit

Regenerate 4% Moonlight every 1s while within 30 units of Luna and not using Moonlight.

Level 2

Broken Deity

Guardian Spirit

Increase Luna's deploy range by 60%.

Level 3

Life Link

Guardian Spirit

[Guardian Spirit] Luna Heals allies around her for 320 per second. Reduce Moonlight's Healing to 135 every 0.15s.

Level 1

Out of Time

Hardening

Reduce the additional Damage Reduction given by Hardening by half. For 1.5s after using Hardening, gain its benefits.

Level 3

Collateral Damage

Form Swap

Gain 30% Lifesteal with Caustic Spray for 3s after swapping into Planted Form.

Level 3

The Great Worm

Form Swap

Gain a 160-Health Shield for 2s after swapping into Travel Form.

Level 2

Raze and Burn

Piercing Quills

Heal for 100 after hitting an enemy with Piercing Quills.

Level 2

Deadly Predator

Weapon

Increase your maximum acid capacity by 40%.

Level 5

Unnatural Persistence

Armor

Heal for 25 every 0.25s while on an objective and for 1s after leaving the objective. This effect increases the longer you maintain it, up to 65 every 0.25s.

Level 1

Feral Strength

Guardian Spirit

Increase Luna's Health by 900.

Level 4

Swift Arrows

Moonlight

Increase allies' Movement Speed by 18% while they are being Healed by Moonlight.

Level 3

Moonwalk

Armor

Increase your maximum Moonlight capacity by 24%.

Level 3

Protectors

Guardian Spirit

Regenerate 4% Moonlight every 1s while within 30 units of Luna and not using Moonlight.

Level 2

Broken Deity

Guardian Spirit

Increase Luna's deploy range by 60%.

Level 3

Sacrifice

Guardian Spirit

[Guardian Spirit] If Io falls below 200 Health while Luna is active, she trades places with Luna, Heals for 50% of her Health, and Luna dies in her stead.

Level 1

Essence Rip

Rend Soul

Increase the Healing done by Rend Soul by 16%.

Level 2

Revenant

Armor

Increase your maximum Health by 200.

Level 4

Ebon Dynamo

Weapon

Increase your maximum Ammo by 4.

Level 2

Dark Sight

Rend Soul

Every Soul Charge you detonate generates 2 Ammo.

Level 2

Umbral Gait

Weapon

Your Movement Speed is increased by 40% while any enemy is marked with a Soul Orb stack.

Level 5

Level 1