Harbinger

Soul Harvest

Reduce the Cooldown of Juggernaut by 1s after activating Soul Harvest.

Level 1

Sinister Allies

Armor

Increase your maximum Health by 450.

Level 3

Hellish Lodestones

Ignition

Reduce your damage taken by 6% for 3s after activating Ignition.

Level 2

Shattered Essence

Soul Harvest

Each Soul Fragment gathered Heals you for 75.

Level 5

Void Lord

Soul Harvest

Reduce your damage taken by 20% for 3s after activating Soul Harvest.

Level 4

Subservience

Soul Harvest

All your living allies are Healed for 1.5% of their maximum Health for each Soul Harvested.

Level 1

Gathering Power

Heal for 70 for each enemy hit with Reckoning.

Level 1

Eternal Strife

Using an ability above Ire threshold Heals you for 300.

Level 4

Overwhelming Presence

Heal for 12 every 0.5s while at least one wall segment of Sanctuary is active.

Level 1

Grim Deliverance

Heal for 25 for every percentage point of Ire consumed by abilities.

Level 5

Tools of Salvation

Increase your maximum Health by 600.

Level 4

Tempering

Augment Reckoning, Sanctuary, and Conviction, but increase their base Cooldown by 25%.

Level 1

Overgrowth

Vine

Reset the Cooldown of Vine after dropping to or below 30% Health.

Level 2

Verdant Expanse

Blossom

Increase the radius of Blossom by 12%.

Level 3

Chop Down

Crippling Throw

Reduce the Cooldown of Crippling Throw by 1.2s.

Level 2

Pick Up

Crippling Throw

Reduce the Cooldown of Vine by 4s after hitting an enemy with Crippling Throw.

Level 4

Fatalis

Crippling Throw

Hitting an enemy with Crippling Throw reduces the Cooldown of Blossom by 2.4s.

Level 4

Rampant Blooming

Vine

Heal for a burst of 500 and an additional 300 over 2s in a 50 unit radius at your location when you finish Vining. Curative (+20% Healing)

Level 1

Fire from Above

Explosive Personality

Increase the vertical movement from each use of Explosive Personality by 40%.

Level 1

Queen's Entrance

Explosive Personality

Increase the lateral movement from each use of Explosive Personality by 40%.

Level 1

Eureka Moment

Weapon

Generate 1 ammo for every -600 explosive damage done.

Level 3

Queen's Reign

Cluster Grenade

Reduce the Cooldown of Cluster Grenade by 0.5s for each enemy hit by the initial and secondary explosions.

Level 5

From Betty with Love

Cluster Grenade

Increase the explosion radius of Cluster Grenade's initial explosion by 30%.

Level 5

Level 1

Fire from Above

Explosive Personality

Increase the vertical movement from each use of Explosive Personality by 40%.

Level 1

Queen's Entrance

Explosive Personality

Increase the lateral movement from each use of Explosive Personality by 40%.

Level 1

Eureka Moment

Weapon

Generate 1 ammo for every -600 explosive damage done.

Level 3

Queen's Reign

Cluster Grenade

Reduce the Cooldown of Cluster Grenade by 0.5s for each enemy hit by the initial and secondary explosions.

Level 5

From Betty with Love

Cluster Grenade

Increase the explosion radius of Cluster Grenade's initial explosion by 30%.

Level 5

Level 1

Guerrilla Tactics

Stealth

Generate 45 Energy after getting an Elimination.

Level 3

Stalker

Stealth

Increase your Reload Speed by 20% while Stealth is active.

Level 2

Grizzled

Armor

Increase your maximum Health by 100.

Level 2

Tactical Retreat

Stealth

Increase your Movement Speed by 40% for 2s after entering Stealth.

Level 4

Roost

Scope

Hits with Talon Rifle generate 4% Ultimate charge.

Level 4

Nocturnal

Stealth

Reduce the rate of Stealth Energy consumption by 50% and increase your Movement Speed by 20% while in Stealth.

Level 1

Violent Birth

Hardening

Reduce the Cooldown of Acceleration by 1.6s after activating Hardening.

Level 2

Triumphant Return

Form Swap

Reduce your acid consumption by 90% for 0.4s after swapping into Planted Form.

Level 1

Deadly Pursuit

Form Swap

Increase your Movement Speed by 20% for 3s after swapping into Travel Form.

Level 3

Towering Presence

Armor

Increase the Health of your Ultimate Health pool by 800.

Level 4

Out of Time

Hardening

Reduce the additional Damage Reduction given by Hardening by half. For 2.5s after using Hardening, gain its benefits.

Level 5

Sight Begets Strength

Primal Vision

Allied players within range of your Primal Vision gain a stackable 90-Health Shield and 15% increased Movement Speed for 4s each time you pulse, but your Primal Vision has its Cooldown increased by 2s.

Level 1

Surf

Shell Spin

Increase the travel distance of Shell Spin by 20%.

Level 2

Ebb and Flow

Armor

Reduce the Cooldown of Shell Spin by 20% after falling to or below 40% Health.

Level 1

Rampage

Armor

Increase your Movement Speed by 40% for 4s after getting an Elimination.

Level 5

Strongarm

Dredge Anchor

Reduce the Cooldown of Dredge Anchor by 3s.

Level 5

Salvage

Dredge Anchor

Regenerate 1 Ammo every 0.5s for 1s after activating Dredge Anchor.

Level 2

Leviathan

Ancient Rage

Increase your Ultimate charge rate by 25% and Health by 500. Activating Ancient Rage resets the Cooldown of Dredge Anchor and Shell Spin, and increases Movement Speed by 20%.

Level 1

Launch

Charge

Increase the travel distance of Charge by 15%.

Level 1

Running Start

Charge

Increase your Movement Speed by 20% for 3s after using Charge.

Level 2

Incinerate

Fireball

Reduce the Cooldown of Fireball by 2.25s.

Level 5

Hot Pursuit

Fireball

Increase your Movement Speed by 32% for 3s after hitting an enemy with Fireball.

Level 4

Looks That Kill

Fireball

Reduce the Cooldown of Charge by 1.7999999999999998s after hitting an enemy with Fireball.

Level 3

Scorch

Fireball

Increase the damage of Fireball by 35%, decrease its Cooldown by 1s, and deal 20% increased damage for each subsequent target hit after the first.

Level 1

Tracker

Hunter's Mark

Increase your Movement Speed by 25% while Hunter's Mark is active.

Level 5

Survivalist

Armor

Increase your maximum Health by 50.

Level 1

Ranger

Hunter's Mark

Reduce the Cooldown of Hunter’s Mark by 4s.

Level 4

Primal Might

Armor

Reduce your active Cooldowns by 30% after getting an Elimination.

Level 3

Favored Quarry

Hunter's Mark

Increase the duration of Hunter's Mark by 1.2s.

Level 2

Burn, Monster!

Fire Bomb

Fire Bomb deals 30% increased damage and applies a 60% reduce healing effect.

Level 1

Relativity

Astral Mark

Heal for 120 every 1s for 10s after applying Astral Mark.

Level 3

Sidereal

Weapon

Increase your maximum Health by 50.

Level 1

Retrograde

Astral Mark

Reduce the Cooldown of Astral Mark by 1.6s.

Level 4

Lightyears

Astral Mark

Increase the range of Astral Mark by 12%.

Level 3

Astral Cycle

Astral Mark

Increase the duration of Astral Mark by 1.6s.

Level 4

Luminary

Astral Mark

All allies already affected by Astral Mark are Healed for an additional 285 when you use Astral Mark.

Level 1

Level 4

Level 1

Level 3

Level 4

Level 3

Level 1

Relativity

Astral Mark

Heal for 120 every 1s for 10s after applying Astral Mark.

Level 3

Sidereal

Weapon

Increase your maximum Health by 50.

Level 1

Retrograde

Astral Mark

Reduce the Cooldown of Astral Mark by 1.6s.

Level 4

Lightyears

Astral Mark

Increase the range of Astral Mark by 12%.

Level 3

Astral Cycle

Astral Mark

Increase the duration of Astral Mark by 1.6s.

Level 4

Luminary

Astral Mark

All allies already affected by Astral Mark are Healed for an additional 285 when you use Astral Mark.

Level 1

Shear

Earthen Guard

Generate 1 Ammo after activating Earthen Guard.

Level 1

Lodestone

Warder's Field

Generate 2 Ammo after activating Warder's Field.

Level 1

Geomancer

Earthen Guard

Reduce the Cooldown of Earthen Guard by 2.5s.

Level 5

Sacred Ground

Warder's Field

Reduce your damage taken by 25% while standing in Warder's Field.

Level 5

Steadfast

Armor

Increase your maximum Health by 450.

Level 3

Mother's Grace

Earthen Guard

Reduce your damage taken by 15%, gain an additional 20% Bonus Healing, and gain Immunity to Crowd Control while Earthen Guard is active.

Level 1