Feral Strength

Guardian Spirit

Increase Luna's Health by 900.

Level 4

Swift Arrows

Moonlight

Increase allies' Movement Speed by 18% while they are being Healed by Moonlight.

Level 3

Restored Faith

Moonlight

Heal for 40 every time you Heal an ally with Moonlight.

Level 4

Sky-Walker

Armor

Lower your gravity and increase your air control by 10% while firing your Light Bow.

Level 1

Full Moon

Lunar Leap

Gain a 300-Health Shield for 3s after activating Lunar Leap.

Level 3

Sacrifice

Guardian Spirit

[Guardian Spirit] If Io falls below 200 Health while Luna is active, she trades places with Luna, Heals for 50% of her Health, and Luna dies in her stead.

Level 1

The Great Worm

Form Swap

Gain a 240-Health Shield for 2s after swapping into Travel Form.

Level 3

Violent Birth

Hardening

Reduce the Cooldown of Acceleration by 1.6s after activating Hardening.

Level 2

Triumphant Return

Form Swap

Reduce your acid consumption by 90% for 0.8s after swapping into Planted Form.

Level 2

Deadly Pursuit

Form Swap

Increase your Movement Speed by 20% for 3s after swapping into Travel Form.

Level 3

Towering Presence

Armor

Increase the Health of your Ultimate Health pool by 1000.

Level 5

Corrosive Acid

Caustic Spray

Augment your acid, increasing the damage done by Caustic Spray to 42, the damage of acid pools to 40, and increase the Movement Speed of allies within your acid pools by 20%.

Level 1

Antediluvian

Overcharge

Increase the duration of Overcharge by 0.8s.

Level 4

Daredevil

Blitz Upper

Heal for 210 after activating Blitz Upper.

Level 3

Primeval Might

Overcharge

Gain 15% Lifesteal while Overcharge is active.

Level 3

Spirited

Armor

Increase your maximum Health by 150

Level 3

Reclamation

Overcharge

Reduce the Cooldown of Overcharge by 1s.

Level 2

Nothing Personal

True Power

Increase your Ultimate charge rate by 15% and enemies at or below 65% Health are Revealed to you.

Level 1

Celestial Body

Armor

Increase your maximum Health by 100.

Level 2

Swift Arrows

Moonlight

Increase allies' Movement Speed by 24% while they are being Healed by Moonlight.

Level 4

Crescent

Lunar Leap

Reset the Cooldown of Lunar Leap after dropping to or below 30% Health.

Level 2

Restored Faith

Moonlight

Heal for 30 every time you Heal an ally with Moonlight.

Level 3

Moonlight Garden

Weapon

Generate 8% Moonlight for each enemy hit with Light Bow.

Level 4

Goddess' Blessing

Moonlight

Allies being Healed by Moonlight take 15% reduced damage.

Level 1

Broken Deity

Guardian Spirit

Increase Luna's deploy range by 20%.

Level 1

Sky-Walker

Armor

Lower your gravity and increase your air control by 30% while firing your Light Bow.

Level 3

Half Moon

Lunar Leap

Reduce the Cooldown of Lunar Leap by 4s.

Level 5

Full Moon

Lunar Leap

Gain a 300-Health Shield for 3s after activating Lunar Leap.

Level 3

Crescent

Lunar Leap

Reset the Cooldown of Lunar Leap after dropping to or below 45% Health.

Level 3

Sacrifice

Guardian Spirit

[Guardian Spirit] If Io falls below 200 Health while Luna is active, she trades places with Luna, Heals for 50% of her Health, and Luna dies in her stead.

Level 1

Trigger Happy

Weapon

Increase your Healing received by 15% while firing your SMGs.

Level 3

Surprise!

Shadow Step

Generate 15 Ammo after activating Shadow Step.

Level 3

Tenacious

Shadow Step

Heal for 160 after activating Shadow Step.

Level 4

Trespasser

Agility

Increase your jump height by 20% while Agility is active.

Level 1

Memento

Weapon

Increase your maximum Ammo by 20.

Level 4

Level 1

Somersault

Armor

Reset the Cooldown of Dodge Roll after dropping to or below 45% Health.

Level 3

Quiver

Weapon

Increase your maximum Ammo by 2.

Level 2

Incitement

Dodge Roll

Reduce the Cooldown of Dodge Roll by 5s if you hit an enemy with your first weapon shot after using Dodge Roll.

Level 5

Lunge

Dodge Roll

Increase the distance traveled when using Dodge Roll by 20%.

Level 2

Onslaught

Dodge Roll

Activating Dodge Roll heals you for 225.

Level 3

Exaction

Dodge Roll

For 8s after using Dodge Roll, your next weapon shot's damage is increased by 30%.

Level 1

Antediluvian

Overcharge

Increase the duration of Overcharge by 0.8s.

Level 4

Daredevil

Blitz Upper

Heal for 210 after activating Blitz Upper.

Level 3

Primeval Might

Overcharge

Gain 15% Lifesteal while Overcharge is active.

Level 3

Spirited

Armor

Increase your maximum Health by 150

Level 3

Reclamation

Overcharge

Reduce the Cooldown of Overcharge by 1s.

Level 2

Nothing Personal

True Power

Increase your Ultimate charge rate by 15% and enemies at or below 65% Health are Revealed to you.

Level 1

Antediluvian

Overcharge

Increase the duration of Overcharge by 0.8s.

Level 4

Daredevil

Blitz Upper

Heal for 210 after activating Blitz Upper.

Level 3

Primeval Might

Overcharge

Gain 15% Lifesteal while Overcharge is active.

Level 3

Spirited

Armor

Increase your maximum Health by 150

Level 3

Reclamation

Overcharge

Reduce the Cooldown of Overcharge by 1s.

Level 2

Nothing Personal

True Power

Increase your Ultimate charge rate by 15% and enemies at or below 65% Health are Revealed to you.

Level 1

Fuel Reserves

Advance

Increase the travel distance of Advance by 35%.

Level 5

Proximity

Advance

Reduce your damage taken by 12% for 3s after activating Advance.

Level 4

Dampener

Armor

Increase your maximum Health by 150.

Level 1

Refraction

Emitter

Reduce the Cooldown of Emitter by 2s.

Level 2

Air-Cooled

Advance

Heal for 240 after activating Advance.

Level 3

Rocket Barrage

Missile Launcher

Increase the radius of Missile Launcher's explosions by 66%.

Level 1