Run and Gun

Hustle

Regenerate 4 Ammo every 1s while using Hustle.

Level 2

Scramble

Hustle

Reduce damage taken by 12% while using Hustle.

Level 4

Hot Potato

Frag Grenade

Reduce the cook time of Frag Grenade by 20%.

Level 2

Second Wind

Hustle

Increase your Movement Speed after deactivating Hustle by 30% for 1.5s.

Level 3

Guerrilla Warfare

Hustle

Reduce the Cooldown of Frag Grenade by 0.8s every 1s while Hustle is active.

Level 4

Shrapnel

Frag Grenade

Increase the radius of Frag Grenade's explosion by 40%.

Level 1

Run and Gun

Hustle

Regenerate 4 Ammo every 1s while using Hustle.

Level 2

Scramble

Hustle

Reduce damage taken by 12% while using Hustle.

Level 4

Hot Potato

Frag Grenade

Reduce the cook time of Frag Grenade by 20%.

Level 2

Second Wind

Hustle

Increase your Movement Speed after deactivating Hustle by 30% for 1.5s.

Level 3

Guerrilla Warfare

Hustle

Reduce the Cooldown of Frag Grenade by 0.8s every 1s while Hustle is active.

Level 4

Burst Mode

Weapon

Switch your rifle to Three-Shot-Burst mode, increasing your effective range and firing three 210-damage shots every 0.4s.

Level 1

Somersault

Armor

Reset the Cooldown of Dodge Roll after dropping to or below 45% Health.

Level 3

Quiver

Weapon

Increase your maximum Ammo by 2.

Level 2

Incitement

Dodge Roll

Reduce the Cooldown of Dodge Roll by 5s if you hit an enemy with your first weapon shot after using Dodge Roll.

Level 5

Lunge

Dodge Roll

Increase the distance traveled when using Dodge Roll by 20%.

Level 2

Onslaught

Dodge Roll

Activating Dodge Roll heals you for 225.

Level 3

Impulse

Blast Shot

Blast Shot has its damage increased by 150 and its Cooldown decreased by 2s.

Level 1

Perennial

Blossom

Reduce the Cooldown of Blossom by 2s.

Level 5

Unstoppable

Blossom

Reduce your damage taken by 24% for 4s after using Blossom.

Level 4

Overgrowth

Vine

Reset the Cooldown of Vine after dropping to or below 30% Health.

Level 2

Heavenly Agility

Vine

Gain a 200 Shield for 3s after using Vine.

Level 2

Vine Grasp

Vine

Increase the range of Vine by 20%.

Level 2

Ferocity

Weapon

Increase your maximum damage scaling over distance by 60%. Surgery (+10% Damage)

Level 1

Depths of Despair

Reduce the damage you take by 12% for 2s after activating Conviction.

Level 3

Tools of Salvation

Increase your maximum Health by 300.

Level 2

Ad Eternum

Reduce the damage you take by 6% while at least one wall segment of Sanctuary is active.

Level 2

Grim Deliverance

Heal for 25 for every percentage point of Ire consumed by abilities.

Level 5

Forged in Battle

Reduce the Cooldown of Sanctuary by 1.7999999999999998s.

Level 3

Persistence

Gain 30% Lifesteal on your hits with Judgement.

Level 1

Refraction

Emitter

Reduce the Cooldown of Emitter by 5s.

Level 5

Metal March

Missile Launcher

Reduce your damage taken by 9% for 2s after using Missile Launcher.

Level 3

Dampener

Armor

Increase your maximum Health by 450.

Level 3

No Chill

Weapon

Increase your Movement Speed while firing by 5%.

Level 1

Warden

Emitter

Increase the duration of Emitter by 3s.

Level 3

Rocket Barrage

Missile Launcher

Increase the radius of Missile Launcher's explosions by 66%.

Level 1

Eternal

Rune of Travel

Increase the duration of Rune of Travel by 1s.

Level 1

Spirited

Armor

Increase your maximum Health by 200

Level 4

Revitalize

Rune of Travel

Heal for 200 after Teleporting to your Rune of Travel.

Level 2

Primeval Might

Overcharge

Gain 20% Lifesteal while Overcharge is active.

Level 4

Guts

Armor

Reduce your damage taken by 20% while at or below 40% Health.

Level 4

Faustian Bargain

Rune of Travel

Rune of Travel no longer automatically activates, and if the Rune is not used the Cooldown is reset.

Level 1

Mirage

Withdraw

Reduce the Cooldown of Withdraw by 3s.

Level 5

Poise

Armor

Reduce the Movement Speed penalty while drawing your Longbow by 60%.

Level 3

Quicksand

Crippling Arrow

Reduce the Cooldown of Crippling Arrow by 1.2s.

Level 2

Skewer

Crippling Arrow

Hitting an enemy with Crippling Arrow reduces the Cooldown of Rapid Shot by 1.2s.

Level 2

Run Like the Wind

Withdraw

Increase your Movement Speed by 21% for 3s after using Withdraw.

Level 3

Desert Silence

Crippling Arrow

Reduce the Cooldown of Crippling Arrow by 1s. Crippling Arrow now Silences and Slows enemies by 15% for 2s instead.

Level 1

Unholy Flight

Death Wings

Reduce your damage taken by 10% for 1.5s after activating Death Wings.

Level 2

Virulent

Blood Hex

Reduce the Blood cost of Death Wings by 4% while an enemy is marked by Blood Hex.

Level 1

Blood Sense

Blood Hex

Hitting an enemy with Blood Hex will Reveal them to your team for 2s.

Level 4

Sanguine Pact

Death Wings

Reduce the Cooldown of Blood Hex by 2s after activating Death Wings.

Level 4

Wings of Terror

Death Wings

Increase the lateral movement of Death Wings by 24%.

Level 4

Cursed Accord

Armor

Decrease the Blood cost of your abilities by 35% and increase your Healing done by 15%, but reduce the damage you do by 25%.

Level 1

Unholy Flight

Death Wings

Reduce your damage taken by 10% for 1.5s after activating Death Wings.

Level 2

Virulent

Blood Hex

Reduce the Blood cost of Death Wings by 4% while an enemy is marked by Blood Hex.

Level 1

Blood Sense

Blood Hex

Hitting an enemy with Blood Hex will Reveal them to your team for 2s.

Level 4

Sanguine Pact

Death Wings

Reduce the Cooldown of Blood Hex by 2s after activating Death Wings.

Level 4

Wings of Terror

Death Wings

Increase the lateral movement of Death Wings by 24%.

Level 4

Cursed Accord

Armor

Decrease the Blood cost of your abilities by 35% and increase your Healing done by 15%, but reduce the damage you do by 25%.

Level 1

Perennial

Blossom

Reduce the Cooldown of Blossom by 2s.

Level 5

Unstoppable

Blossom

Reduce your damage taken by 24% for 4s after using Blossom.

Level 4

Overgrowth

Vine

Reset the Cooldown of Vine after dropping to or below 30% Health.

Level 2

Heavenly Agility

Vine

Gain a 200 Shield for 3s after using Vine.

Level 2

Vine Grasp

Vine

Increase the range of Vine by 20%.

Level 2

Ferocity

Weapon

Increase your maximum damage scaling over distance by 60%. Surgery (+10% Damage)

Level 1

Gift-Giver

Healing Potion

Heal yourself for 60% of Healing Potion's effect if you hit an ally but not yourself.

Level 3

Sprint

Weightless

Increase your jump speed by an additonal 20% while using Weightless.

Level 2

Reload

Healing Potion

Reduce the Cooldown of Healing Potion by 3s for each ally it hits.

Level 5

Refreshing Jog

Weightless

Heal for 60 every 1s while Weightless is active.

Level 1

Medicinal Excellence

Healing Potion

Increase the radius of Healing Potion by 32%.

Level 4

Mega Potion

Healing Potion

Healing Potion's Healing is increased by 100% and now provides a Stacking 200 Shield. Remedy (+5% Damage and +10% Healing Done)

Level 1

Guerrilla Tactics

Stealth

Generate 45 Energy after getting an Elimination.

Level 3

Stalker

Stealth

Increase your Reload Speed by 20% while Stealth is active.

Level 2

Grizzled

Armor

Increase your maximum Health by 100.

Level 2

Tactical Retreat

Stealth

Increase your Movement Speed by 40% for 2s after entering Stealth.

Level 4

Roost

Scope

Hits with Talon Rifle generate 4% Ultimate charge.

Level 4

Nocturnal

Stealth

Reduce the rate of Stealth Energy consumption by 50% and increase your Movement Speed by 20% while in Stealth.

Level 1

Lifegiver

Protection

Heal the target of your Protection for 300.

Level 4

Eldritch Speed

Recharge

Increase your Movement Speed by 20% during Recharge and for 3s after it ends.

Level 2

Winddancer

Protection

Increase the Movement Speed of your Protection target by 40% for 2s.

Level 4

Hearthwarder

Protection

Generate 225 Shield after using Protection.

Level 3

Vital Grasp

Recharge

Heal for 150 after activating Recharge.

Level 2

Direct Current

Nullify

Double the range at which you can use Nullify.

Level 1