Perennial

Blossom

Reduce the Cooldown of Blossom by 1.6s.

Level 4

Fatalis

Crippling Throw

Hitting an enemy with Crippling Throw reduces the Cooldown of Blossom by 1.2s.

Level 2

Force of Nature

Armor

Gain a 300-Health Shield for 3s after dropping to or below 50% Health.

Level 2

Gentle Breeze

Blossom

Increase your allies' Movement Speed by 15% for 4s after they are Healed by Blossom.

Level 3

Verdant Expanse

Blossom

Increase the radius of Blossom by 16%.

Level 4

Rampant Blooming

Vine

Heal for a burst of 500 and an additional 300 over 2s in a 50 unit radius at your location when you finish Vining. Curative (+20% Healing)

Level 1

Healing Station

Turret

Heal for 75 every 1s while standing near any of your Turrets.

Level 3

Combat Repair

Turret

Heal all of your Turrets for 270 every 1s while you are standing near them.

Level 3

Double-Time

Rocket Boots

Heal for 100 every 1s for 3s after activating Rocket Boots.

Level 2

Bowling Ball

Rocket Boots

Gain a 300-Health Shield for 3s after activating Rocket Boots.

Level 2

Bunker

Barricade

Increase the Health of Barricade by 1000.

Level 5

Tinkerin

Weapon

Modify your Blunderbuss to fire a single slug that deals 600 damage.

Level 1

Healing Station

Turret

Heal for 75 every 1s while standing near any of your Turrets.

Level 3

Combat Repair

Turret

Heal all of your Turrets for 270 every 1s while you are standing near them.

Level 3

Double-Time

Rocket Boots

Heal for 100 every 1s for 3s after activating Rocket Boots.

Level 2

Bowling Ball

Rocket Boots

Gain a 300-Health Shield for 3s after activating Rocket Boots.

Level 2

Bunker

Barricade

Increase the Health of Barricade by 1000.

Level 5

Tinkerin

Weapon

Modify your Blunderbuss to fire a single slug that deals 600 damage.

Level 1

Chase

Prowl

Increase the Movement Speed bonus while using Prowl by 30%.

Level 3

Walk it Off

Prowl

Heal for 160 every 1s while Prowl is active.

Level 4

Predation

Armor

Reduce the Cooldown of Prowl by 0.6000000000000001s each time you hit an enemy with a dagger.

Level 3

Persistence

Armor

Reduce the damage you take by 16% while at or below 40% Health.

Level 4

On Edge

Nine Lives

Reduce the Cooldown of Nine Lives by 1s.

Level 1

Cat Burglar

Prowl

Increase the damage of each of your daggers from your first dagger sets within 5s of Prowl ending by 30%.

Level 1

Astral Cycle

Astral Mark

Increase the duration of Astral Mark by 1.6s.

Level 4

Retrograde

Astral Mark

Reduce the Cooldown of Astral Mark by 2s.

Level 5

Relativity

Astral Mark

Heal for 120 every 1s for 10s after applying Astral Mark.

Level 3

Lightyears

Astral Mark

Increase the range of Astral Mark by 8%.

Level 2

Cosmic Barrier

Stellar Wind

Reduce your damage taken by 5% while Stellar Wind is active.

Level 1

Luminary

Astral Mark

All allies already affected by Astral Mark are Healed for an additional 285 when you use Astral Mark.

Level 1

Perennial

Blossom

Reduce the Cooldown of Blossom by 1.6s.

Level 4

Fatalis

Crippling Throw

Hitting an enemy with Crippling Throw reduces the Cooldown of Blossom by 1.2s.

Level 2

Force of Nature

Armor

Gain a 300-Health Shield for 3s after dropping to or below 50% Health.

Level 2

Gentle Breeze

Blossom

Increase your allies' Movement Speed by 15% for 4s after they are Healed by Blossom.

Level 3

Verdant Expanse

Blossom

Increase the radius of Blossom by 16%.

Level 4

Rampant Blooming

Vine

Heal for a burst of 500 and an additional 300 over 2s in a 50 unit radius at your location when you finish Vining. Curative (+20% Healing)

Level 1

Healing Station

Turret

Heal for 75 every 1s while standing near any of your Turrets.

Level 3

Combat Repair

Turret

Heal all of your Turrets for 270 every 1s while you are standing near them.

Level 3

Double-Time

Rocket Boots

Heal for 100 every 1s for 3s after activating Rocket Boots.

Level 2

Bowling Ball

Rocket Boots

Gain a 300-Health Shield for 3s after activating Rocket Boots.

Level 2

Bunker

Barricade

Increase the Health of Barricade by 1000.

Level 5

Forgefire

Dome Shield

Dome Shield now costs 40% Ultimate charge and places a Flamethrower Turret with modified values and no Shield. Can have two active at once.

Level 1

Restored Faith

Moonlight

Heal for 30 every time you Heal an ally with Moonlight.

Level 3

Moonlight Garden

Weapon

Generate 6% Moonlight for each enemy hit with Light Bow.

Level 3

Moonwalk

Armor

Increase your maximum Moonlight capacity by 40%.

Level 5

Crescent

Lunar Leap

Reset the Cooldown of Lunar Leap after dropping to or below 30% Health.

Level 2

Swift Arrows

Moonlight

Increase allies' Movement Speed by 12% while they are being Healed by Moonlight.

Level 2

Sacrifice

Guardian Spirit

[Guardian Spirit] If Io falls below 200 Health while Luna is active, she trades places with Luna, Heals for 50% of her Health, and Luna dies in her stead.

Level 1

Overwhelming Presence

Heal for 36 every 0.5s while at least one wall segment of Sanctuary is active.

Level 3

Ad Eternum

Reduce the damage you take by 9% while at least one wall segment of Sanctuary is active.

Level 3

Grim Deliverance

Heal for 15 for every percentage point of Ire consumed by abilities.

Level 3

Depths of Despair

Reduce the damage you take by 12% for 2s after activating Conviction.

Level 3

Solemn Watch

Increase the distance traveled by Conviction by 18%.

Level 3

Persistence

Gain 30% Lifesteal on your hits with Judgement.

Level 1

Swift Arrows

Moonlight

Increase allies' Movement Speed by 12% while they are being Healed by Moonlight.

Level 2

Moonlight Garden

Weapon

Generate 6% Moonlight for each enemy hit with Light Bow.

Level 3

Moonwalk

Armor

Increase your maximum Moonlight capacity by 40%.

Level 5

Spirit Arrows

Moonlight

Reduce the Cooldown of Luna's redeploy by 0.2s after healing with Moonlight.

Level 2

Restored Faith

Moonlight

Heal for 30 every time you Heal an ally with Moonlight.

Level 3

Sacrifice

Guardian Spirit

[Guardian Spirit] If Io falls below 200 Health while Luna is active, she trades places with Luna, Heals for 50% of her Health, and Luna dies in her stead.

Level 1

The Hunted

Retribution

You gain 15% Lifesteal against your Retribution target.

Level 3

Compel

In Pursuit

Reduce the Cooldown of In Pursuit by 3.5999999999999996s.

Level 3

Hardiness

In Pursuit

Decrease your damage taken by 12% while using In Pursuit.

Level 2

Conditioned

Armor

Increase your Movement Speed by 12%.

Level 4

Justice Served

Retribution

Heal for 45% of your maximum Health after getting a Killing Blow on your Retribution target.

Level 3

Discovery

Retribution

Increase the damage you deal to your Retribution target by 25%.

Level 1

Deadly Predator

Weapon

Increase your maximum acid capacity by 32%.

Level 4

Raze and Burn

Piercing Quills

Heal for 150 after hitting an enemy with Piercing Quills.

Level 3

Last Efforts

Piercing Quills

Hitting an enemy with Piercing Quills grants 20% Lifesteal for 1.7999999999999998s.

Level 3

Dreadful Compact

Piercing Quills

Reduce the Cooldown of Hardening by 0.8999999999999999s after hitting an enemy with Piercing Quills.

Level 3

Ancient Wounds

Piercing Quills

Generate 4% of your maximum acid after hitting an enemy with Piercing Quills.

Level 2

Unnatural Persistence

Armor

Heal for 25 every 0.25s while on an objective and for 1s after leaving the objective. This effect increases the longer you maintain it, up to 65 every 0.25s.

Level 1

True Grit

Sniper Mode

Hitting an enemy with a fully-charged Sniper Mode shot Heals you for 200.

Level 4

Beam Me Up

Transporter

Increase the Flight Speed of Transporter by 40%.

Level 4

Amplitude

Oppressor Mine

Increase the range of Oppressor Mine beams by 60%.

Level 3

Open Season

Sniper Mode

Hitting an enemy with a fully-charged Sniper Mode shot generates 1 Ammo.

Level 1

Quick Scope

Sniper Mode

Reduce scope-in and scope-out time by 60%.

Level 3

Level 1

Swift Arrows

Moonlight

Increase allies' Movement Speed by 12% while they are being Healed by Moonlight.

Level 2

Moonlight Garden

Weapon

Generate 6% Moonlight for each enemy hit with Light Bow.

Level 3

Moonwalk

Armor

Increase your maximum Moonlight capacity by 40%.

Level 5

Spirit Arrows

Moonlight

Reduce the Cooldown of Luna's redeploy by 0.2s after healing with Moonlight.

Level 2

Restored Faith

Moonlight

Heal for 30 every time you Heal an ally with Moonlight.

Level 3

Sacrifice

Guardian Spirit

[Guardian Spirit] If Io falls below 200 Health while Luna is active, she trades places with Luna, Heals for 50% of her Health, and Luna dies in her stead.

Level 1

Haunting

Ghost Walk

Reduce the Cooldown of Ghost Walk by 1.2s.

Level 2

Gale

Ghost Walk

Decrease the Cooldown of Healing Totem by 1.2s after activating Ghost Walk.

Level 2

Phantom

Ghost Walk

Heal for 100 over the duration of Ghost Walk.

Level 1

Electrostatic

Weapon

Increase your Ammo regeneration speed by 60%.

Level 5

Thunderlord

Weapon

Increase your maximum Ammo by 50%.

Level 5

Spirit's Domain

Weapon

Hitting allies with your Lightning Staff Heals them for 800 per second and your maximum Ammo capacity is increased by 15%. Curative (+20% Healing)

Level 1

Bragging Rights

Weapon

Hitting enemies with Bladed Chakrams Heals you for 75.

Level 5

Honed Senses

Combat Trance

Generate 1.2% Ultimate charge every 1s while Combat Trance is active.

Level 2

Instrument of Fate

Heavy Blade

Reduce the Cooldown of Combat Trance by 1.2000000000000002s for each enemy hit by Heavy Blade.

Level 3

World-traveler

Crouching Tigron

Reset the Cooldown of Crouching Tigron after falling to or below 30% Health.

Level 2

Flying Chakrams

Weapon

Increase your maximum Ammo by 3.

Level 3

Predatory Instincts

Weapon

Bladed Chakrams are able to bounce 2 additional times before disappearing.

Level 1

Bragging Rights

Weapon

Hitting enemies with Bladed Chakrams Heals you for 75.

Level 5

Crowd-Pleaser

Combat Trance

Generate 3 Ammo after activating Combat Trance.

Level 3

Instrument of Fate

Heavy Blade

Reduce the Cooldown of Combat Trance by 0.8s for each enemy hit by Heavy Blade.

Level 2

World-traveler

Crouching Tigron

Reset the Cooldown of Crouching Tigron after falling to or below 30% Health.

Level 2

Flying Chakrams

Weapon

Increase your maximum Ammo by 3.

Level 3

Predatory Instincts

Weapon

Bladed Chakrams are able to bounce 2 additional times before disappearing.

Level 1

Carry On

Illusion

Increase the duration of your Illusions by 2s.

Level 2

Squadron

Illusion

Increase the Health of each of your Illusions by 100.

Level 1

Tangible

Armor

Increase your maximum Health by 200.

Level 4

Shuffle

Dimensional Link

Increase your Movement Speed by 30% while Dimensional Link is active.

Level 3

Brittle

Shatter

Reduce the Cooldown of Shatter by 2.5s.

Level 5

Life Exchange

Shatter

Shatter instantly Heals your target for 800, but its Cooldown is increased by 1s and it no longer explodes your Illusions. Reduce the Cooldown of Shatter by 2s if you miss.

Level 1