Run and Gun

Hustle

Regenerate 4 Ammo every 1s while using Hustle.

Level 2

Scramble

Hustle

Reduce damage taken by 12% while using Hustle.

Level 4

Hot Potato

Frag Grenade

Reduce the cook time of Frag Grenade by 20%.

Level 2

Second Wind

Hustle

Increase your Movement Speed after deactivating Hustle by 30% for 1.5s.

Level 3

Guerrilla Warfare

Hustle

Reduce the Cooldown of Frag Grenade by 0.8s every 1s while Hustle is active.

Level 4

Cardio

Hustle

Heal for 300 per second while using Hustle.

Level 1

Run and Gun

Hustle

Regenerate 4 Ammo every 1s while using Hustle.

Level 2

Scramble

Hustle

Reduce damage taken by 12% while using Hustle.

Level 4

Hot Potato

Frag Grenade

Reduce the cook time of Frag Grenade by 20%.

Level 2

Second Wind

Hustle

Increase your Movement Speed after deactivating Hustle by 30% for 1.5s.

Level 3

Guerrilla Warfare

Hustle

Reduce the Cooldown of Frag Grenade by 0.8s every 1s while Hustle is active.

Level 4

Cardio

Hustle

Heal for 300 per second while using Hustle.

Level 1

Napalm

Fire Bomb

Activating Fire Bomb heals you for 300 over 2s.

Level 3

Volatile

Fire Bomb

Reduce the Cooldown of Fire Bomb by 2.4s.

Level 4

Survivalist

Armor

Increase your maximum Health by 50.

Level 1

Primal Might

Armor

Reduce your active Cooldowns by 30% after getting an Elimination.

Level 3

Incensed

Fire Bomb

Increase your Movement Speed by 32% for 4s after using Fire Bomb.

Level 4

Burn, Monster!

Fire Bomb

Fire Bomb deals 30% increased damage and applies a 60% reduce healing effect.

Level 1

Somersault

Armor

Reset the Cooldown of Dodge Roll after dropping to or below 45% Health.

Level 3

Quiver

Weapon

Increase your maximum Ammo by 2.

Level 2

Incitement

Dodge Roll

Reduce the Cooldown of Dodge Roll by 5s if you hit an enemy with your first weapon shot after using Dodge Roll.

Level 5

Lunge

Dodge Roll

Increase the distance traveled when using Dodge Roll by 20%.

Level 2

Onslaught

Dodge Roll

Activating Dodge Roll heals you for 225.

Level 3

Exaction

Dodge Roll

For 8s after using Dodge Roll, your next weapon shot's damage is increased by 30%.

Level 1

Retrograde

Astral Mark

Reduce the Cooldown of Astral Mark by 2s.

Level 5

Astral Cycle

Astral Mark

Increase the duration of Astral Mark by 2s.

Level 5

Relativity

Astral Mark

Heal for 80 every 1s for 10s after applying Astral Mark.

Level 2

Cosmic Barrier

Stellar Wind

Reduce your damage taken by 5% while Stellar Wind is active.

Level 1

Lightyears

Astral Mark

Increase the range of Astral Mark by 8%.

Level 2

Luminary

Astral Mark

All allies already affected by Astral Mark are Healed for an additional 285 when you use Astral Mark.

Level 1

Run and Gun

Hustle

Regenerate 4 Ammo every 1s while using Hustle.

Level 2

Scramble

Hustle

Reduce damage taken by 12% while using Hustle.

Level 4

Hot Potato

Frag Grenade

Reduce the cook time of Frag Grenade by 20%.

Level 2

Second Wind

Hustle

Increase your Movement Speed after deactivating Hustle by 30% for 1.5s.

Level 3

Guerrilla Warfare

Hustle

Reduce the Cooldown of Frag Grenade by 0.8s every 1s while Hustle is active.

Level 4

Cardio

Hustle

Heal for 300 per second while using Hustle.

Level 1

Broken Deity

Guardian Spirit

Increase Luna's deploy range by 20%.

Level 1

Sky-Walker

Armor

Lower your gravity and increase your air control by 30% while firing your Light Bow.

Level 3

Half Moon

Lunar Leap

Reduce the Cooldown of Lunar Leap by 4s.

Level 5

Full Moon

Lunar Leap

Gain a 300-Health Shield for 3s after activating Lunar Leap.

Level 3

Crescent

Lunar Leap

Reset the Cooldown of Lunar Leap after dropping to or below 45% Health.

Level 3

Sacrifice

Guardian Spirit

[Guardian Spirit] If Io falls below 200 Health while Luna is active, she trades places with Luna, Heals for 50% of her Health, and Luna dies in her stead.

Level 1

A Runaway

Hail of Bombs

Increase the explosion radius of Hail of Bombs' explosives by 30%.

Level 3

Queen's Reign

Cluster Grenade

Reduce the Cooldown of Cluster Grenade by 0.4s for each enemy hit by the initial and secondary explosions.

Level 4

Eureka Moment

Weapon

Generate 1 ammo for every -400 explosive damage done.

Level 2

From Betty with Love

Cluster Grenade

Increase the explosion radius of Cluster Grenade's initial explosion by 12%.

Level 2

Bouncing Betty

Explosive Personality

Heal for 120 for each use of Explosive Personality.

Level 4

Level 1

Hulking Monstrosity

Armor

Increase your maximum Health by 450.

Level 3

Powerslave

Power Siphon

Reduce the Movement Speed penalty during Power Siphon by 15%.

Level 1

Strength of Stone

Calamity Blast

Reduce your damage taken by up to 16% based on the number of Calamity Blast charges you have stored.

Level 4

It Follows

Calamity Blast

Increase your Movement Speed by 10% for each Calamity Charge you have.

Level 5

It Watches

Armor

Generate 6% Power Siphon charge for each enemy hit with Massacre Axe.

Level 2

Undying

Armor

Reduce the damage you take by 20% while at or below 50% Health.

Level 1

Bully

Net Shot

Gain 36% Lifesteal against targets Slowed by Net Shot.

Level 3

Tangled

Net Shot

Increase the duration of Net Shot's Slow by 0.6s.

Level 2

Leg Day

Heroic Leap

Increase the jump strength of Heroic Leap by 10%.

Level 2

Gotcha

Net Shot

Increase your Movement Speed by 40% for 3s after using Net Shot.

Level 4

Concussion

Net Shot

Reduce the Cooldown of Net Shot by 4s.

Level 4

Ensnare

Net Shot

Increase the damage you deal to targets Slowed by Net Shot by 20% and increases the duration of Net Shot's Slow by 0.5s. The damage increase lasts a minimum of 2s.

Level 1

A Runaway

Hail of Bombs

Increase the explosion radius of Hail of Bombs' explosives by 30%.

Level 3

Queen's Reign

Cluster Grenade

Reduce the Cooldown of Cluster Grenade by 0.4s for each enemy hit by the initial and secondary explosions.

Level 4

Eureka Moment

Weapon

Generate 1 ammo for every -400 explosive damage done.

Level 2

From Betty with Love

Cluster Grenade

Increase the explosion radius of Cluster Grenade's initial explosion by 12%.

Level 2

Bouncing Betty

Explosive Personality

Heal for 120 for each use of Explosive Personality.

Level 4

Level 1

Trigger Happy

Weapon

Increase your Healing received by 15% while firing your SMGs.

Level 3

Surprise!

Shadow Step

Generate 15 Ammo after activating Shadow Step.

Level 3

Tenacious

Shadow Step

Heal for 160 after activating Shadow Step.

Level 4

Something to Prove

Armor

Generate 10% Energy after falling to or below 50% Health.

Level 1

Memento

Weapon

Increase your maximum Ammo by 20.

Level 4

Level 1

Controlling Nature

Precision Sights

Reduce the Cooldown of Sensor Drone by 0.4s for every enemy hit while Precision Sights is active.

Level 5

One Step Ahead

Armor

Increase your Movement Speed by 12%.

Level 3

Numbed Heart

Armor

Increase your maximum Health by 100.

Level 2

Nowhere to Hide

Sensor Drone

Sensor Drone's Reveal persists for 1.4s after an enemy leaves its Reveal range.

Level 2

Informants

Sensor Drone

Heal for 255 after an enemy that is Revealed by your Sensor Drone dies.

Level 3

Booby Trap

Sensor Drone

Your Sensor Drones explode, dealing 600 damage and applying a Knockback to those hit when an enemy is within 20 units.

Level 1