Playing God

Power Siphon

Heal for 20% of the damage Absorbed by Power Siphon.

Level 4

Abomination

Calamity Blast

Generate 4% Power Siphon charge for each enemy hit with Calamity Blast.

Level 2

Devastation

Calamity Blast

Heal for 120 for each enemy hit by Calamity Blast.

Level 2

Hulking Monstrosity

Armor

Increase your maximum Health by 750.

Level 5

Necromantic Might

Power Siphon

Reduce the damage required to generate a Calamity Blast charge by 200.

Level 2

Undying

Armor

Reduce the damage you take by 20% while at or below 50% Health.

Level 1

Elemental Barrier

Elemental Shift

Heal for 200 after using Elemental Shift.

Level 4

Affinity

Elemental Shift

Reduce your damage taken by 5% for 2s after activating Elemental Shift.

Level 1

Mystic Fire

Frostfire Glide

Regenerate 5% Mana every 1s while using FrostFire Glide.

Level 1

Elemental Grace

Armor

Gain a 500-health Shield for 3s after dropping to or below 50% Health.

Level 5

Ancient Duty

Armor

Increase your maximum Health by 200.

Level 4

Pyromania

Pyre Ball

Each of your Pyre Balls explodes in a 20-unit-radius area on hit and generate more Mana, but the damage of Pyre Ball when fully charged is reduced to 800.

Level 1

Gloomy Nook

Armor

Gain a 750 Shield for 1.75s after falling to or below 75% Health. Can only happen once every 35s.

Level 5

Waning Moon

Blood Hex

Reduce the Blood cost of Swarm by 5% while an ally is marked by Blood Hex.

Level 1

Overflowing Delights

Swarm

Increase the Radius of Swarm by 8%, but increase its cost by {scale=3|3}%.

Level 4

Blood Cannon

Armor

Increase extra Blood Health by 600 while decreasing base Health by {scale=125|125}.

Level 3

Enriched Blood

Swarm

Increase the damage and Healing of Swarm by 8%, but reduce its duration by {scale=0.35|0.35}s.

Level 2

Level 1

Feral Strength

Guardian Spirit

Increase Luna's Health by 1125.

Level 5

Moonwalk

Armor

Increase your maximum Moonlight capacity by 24%.

Level 3

Sky-Walker

Armor

Lower your gravity and increase your air control by 10% while firing your Light Bow.

Level 1

Restored Faith

Moonlight

Heal for 30 every time you Heal an ally with Moonlight.

Level 3

Full Moon

Lunar Leap

Gain a 300-Health Shield for 3s after activating Lunar Leap.

Level 3

Sacrifice

Guardian Spirit

[Guardian Spirit] If Io falls below 200 Health while Luna is active, she trades places with Luna, Heals for 50% of her Health, and Luna dies in her stead.

Level 1

Shattered Essence

Soul Harvest

Each Soul Fragment gathered Heals you for 45.

Level 3

Infernal Reload

Weapon

Increase your Reload Speed by 21%.

Level 3

Triumphant Ascension

Soul Harvest

Heal for 200 after activating Soul Harvest.

Level 2

Abyssal Connections

Ignition

Gain a 200-Health Shield for 3s after activating Ignition.

Level 2

Sinister Allies

Armor

Increase your maximum Health by 750.

Level 5

Earthsplitter

Cataclysm

Increase your Ultimate charge rate by 60%.

Level 1

Shattered Essence

Soul Harvest

Each Soul Fragment gathered Heals you for 45.

Level 3

Infernal Reload

Weapon

Increase your Reload Speed by 21%.

Level 3

Hellish Lodestones

Ignition

Reduce your damage taken by 6% for 3s after activating Ignition.

Level 2

Abyssal Connections

Ignition

Gain a 200-Health Shield for 3s after activating Ignition.

Level 2

Sinister Allies

Armor

Increase your maximum Health by 750.

Level 5

Earthsplitter

Cataclysm

Increase your Ultimate charge rate by 60%.

Level 1

Feral Strength

Guardian Spirit

Increase Luna's Health by 1125.

Level 5

Moonwalk

Armor

Increase your maximum Moonlight capacity by 24%.

Level 3

Sky-Walker

Armor

Lower your gravity and increase your air control by 10% while firing your Light Bow.

Level 1

Restored Faith

Moonlight

Heal for 30 every time you Heal an ally with Moonlight.

Level 3

Full Moon

Lunar Leap

Gain a 300-Health Shield for 3s after activating Lunar Leap.

Level 3

Sacrifice

Guardian Spirit

[Guardian Spirit] If Io falls below 200 Health while Luna is active, she trades places with Luna, Heals for 50% of her Health, and Luna dies in her stead.

Level 1

Towering Presence

Armor

Increase the Health of your Ultimate Health pool by 400.

Level 2

End of an Era

Hardening

Increase the additional Damage Reduction benefit of Hardening by 20% but reduce its duration by {scale=10|10}%.

Level 1

Triumphant Return

Form Swap

Reduce your acid consumption by 90% for 1.6s after swapping into Planted Form.

Level 4

Deadly Predator

Weapon

Increase your maximum acid capacity by 40%.

Level 5

Level 3

Unnatural Persistence

Armor

Heal for 25 every 0.25s while on an objective and for 1s after leaving the objective. This effect increases the longer you maintain it, up to 65 every 0.25s.

Level 1

Hull Piercer

Harpoon

Reduce the Cooldown of Harpoon by 2.4s.

Level 2

Dark Bargain

Shortcut

Heal for 150 every 1s while standing near a Shortcut.

Level 3

Dreadnought

Armor

Increase your maximum Health by 200.

Level 4

Hangman's Ire

Armor

Gain a 750-Health Shield after dropping to or below 50% Health.

Level 5

Expansive Vault

Shortcut

Jump height is increased for your first jump within 1s after exiting a Shortcut.

Level 1

Hurl

Harpoon

Harpoon no longer has a Cooldown, but no longer Slows enemies hit.

Level 1

Shattered Essence

Soul Harvest

Each Soul Fragment gathered Heals you for 45.

Level 3

Infernal Reload

Weapon

Increase your Reload Speed by 21%.

Level 3

Tormented Fissure

Ignition

Generate 50 Ammo after activating Ignition.

Level 2

Abyssal Connections

Ignition

Gain a 200-Health Shield for 3s after activating Ignition.

Level 2

Sinister Allies

Armor

Increase your maximum Health by 750.

Level 5

Earthsplitter

Cataclysm

Increase your Ultimate charge rate by 60%.

Level 1

Flicker

Blink

Heal for 90 after using Blink.

Level 3

Cold Acclimation

Weapon

Your weapon deals 30% less Self-Damage and no longer applies a Knockback to yourself.

Level 3

Frigid Field

Ice Block

Heal for 240 every 1s while Ice Block is active.

Level 4

Keep Moving

Armor

Increase maximum Health by 90.

Level 3

Swift Witch

Soar

Increase your Movement Speed during Soar by 14%.

Level 2

Over the Moon

Soar

Increase the damage you deal with your weapon shots by 25% for 3s after Soar ends.

Level 1

Doomsday

Grumpy Bomb

Heal for 260 after using Grumpy Bomb.

Level 2

Bomb Shelter

Weapon

Reduce the damage you deal to yourself with Sticky Bombs by 100%.

Level 5

King's Court

Detonate

Heal for 45 for each Sticky Bomb you Detonate.

Level 3

Reinforced Casing

Armor

Increase your maximum Ammo by 3.

Level 3

Explosive Entrance

Poppy Bomb

Heal for 150 after using Poppy Bomb.

Level 2

Chain Reaction

Detonate

Hitting with more than one Sticky Bomb on a use of Detonate increases the damage of each explosion after the first by 30% of its base damage, up to 150%.

Level 1