Heal for 30 every time you Heal an ally with Moonlight.
Level 3
Moonwalk
Armor
Increase your maximum Moonlight capacity by 32%.
Level 4
Sanctum of Faith
Moonlight
Regenerate 5% Moonlight every 1s while not using Moonlight.
Level 5
Crescent
Lunar Leap
Reset the Cooldown of Lunar Leap after dropping to or below 30% Health.
Level 2
Sky-Walker
Armor
Lower your gravity and increase your air control by 10% while firing your Light Bow.
Level 1
Sacrifice
Guardian Spirit
[Guardian Spirit] If Io falls below 200 Health while Luna is active, she trades places with Luna, Heals for 50% of her Health, and Luna dies in her stead.