Level 2

Dreadful Compact

Piercing Quills

Reduce the Cooldown of Hardening by 0.3s after hitting an enemy with Piercing Quills.

Level 1

The Great Worm

Form Swap

Gain a 160-Health Shield for 2s after swapping into Travel Form.

Level 2

End of an Era

Hardening

Increase the additional Damage Reduction benefit of Hardening by 100% but reduce its duration by {scale=10|10}%.

Level 5

Spreading Domain

Hardening

Heal for 500 if you do not cancel Hardening before it ends.

Level 5

Unnatural Persistence

Armor

Heal for 25 every 0.25s while on an objective and for 1s after leaving the objective. This effect increases the longer you maintain it, up to 65 every 0.25s.

Level 1

End of an Era

Hardening

Increase the additional Damage Reduction benefit of Hardening by 100% but reduce its duration by {scale=10|10}%.

Level 5

Dreadful Compact

Piercing Quills

Reduce the Cooldown of Hardening by 0.3s after hitting an enemy with Piercing Quills.

Level 1

Last Efforts

Piercing Quills

Hitting an enemy with Piercing Quills grants 20% Lifesteal for 0.6s.

Level 1

Towering Presence

Armor

Increase the Health of your Ultimate Health pool by 600.

Level 3

Spreading Domain

Hardening

Heal for 500 if you do not cancel Hardening before it ends.

Level 5

Corrosive Acid

Caustic Spray

Augment your acid, increasing the damage done by Caustic Spray to 42, the damage of acid pools to 40, and increase the Movement Speed of allies within your acid pools by 20%.

Level 1

End of an Era

Hardening

Increase the additional Damage Reduction benefit of Hardening by 100% but reduce its duration by {scale=10|10}%.

Level 5

Dreadful Compact

Piercing Quills

Reduce the Cooldown of Hardening by 0.3s after hitting an enemy with Piercing Quills.

Level 1

Last Efforts

Piercing Quills

Hitting an enemy with Piercing Quills grants 20% Lifesteal for 0.6s.

Level 1

Towering Presence

Armor

Increase the Health of your Ultimate Health pool by 600.

Level 3

Spreading Domain

Hardening

Heal for 500 if you do not cancel Hardening before it ends.

Level 5

Sight Begets Strength

Primal Vision

Allied players within range of your Primal Vision gain a stackable 90-Health Shield and 15% increased Movement Speed for 4s each time you pulse, but your Primal Vision has its Cooldown increased by 2s.

Level 1

Celestial Body

Armor

Increase your maximum Health by 100.

Level 2

Swift Arrows

Moonlight

Increase allies' Movement Speed by 24% while they are being Healed by Moonlight.

Level 4

Crescent

Lunar Leap

Reset the Cooldown of Lunar Leap after dropping to or below 30% Health.

Level 2

Restored Faith

Moonlight

Heal for 30 every time you Heal an ally with Moonlight.

Level 3

Moonlight Garden

Weapon

Generate 8% Moonlight for each enemy hit with Light Bow.

Level 4

Life Link

Guardian Spirit

[Guardian Spirit] Luna Heals allies around her for 320 per second. Reduce Moonlight's Healing to 135 every 0.15s.

Level 1

End of an Era

Hardening

Increase the additional Damage Reduction benefit of Hardening by 100% but reduce its duration by {scale=10|10}%.

Level 5

Dreadful Compact

Piercing Quills

Reduce the Cooldown of Hardening by 0.3s after hitting an enemy with Piercing Quills.

Level 1

Last Efforts

Piercing Quills

Hitting an enemy with Piercing Quills grants 20% Lifesteal for 0.6s.

Level 1

Towering Presence

Armor

Increase the Health of your Ultimate Health pool by 600.

Level 3

Spreading Domain

Hardening

Heal for 500 if you do not cancel Hardening before it ends.

Level 5

Sight Begets Strength

Primal Vision

Allied players within range of your Primal Vision gain a stackable 90-Health Shield and 15% increased Movement Speed for 4s each time you pulse, but your Primal Vision has its Cooldown increased by 2s.

Level 1